32 
PACIFIC SCIENCE, Vol. VI, January, 1952 
tube 28-35 mm. long, glabrous outside, but 
hirsutulous within from level of tip of calyx 
lobes to the throat, on proximal side cleft 
about halfway, and from the sinus to the base 
having a distinct suture along which the 
corolla may split in age; corolla lobes 6-15 
mm. long, the outside glabrous, the inside 
hirsutulous except on the hyaline winged 
margins; filaments glabrous, about 33 mm. 
long; anthers 5 mm. long, linear; style 42-48 
mm. long, exserted, stout, glabrous; indu- 
sium 3-4 mm. in diameter, ciliolate on the 
rim; drupe 16-20 mm. long, broadly oval, 
shining black; endocarp 9-13 mm. long, 
8-9 mm. wide, 6 mm. thick, definitely 
flattened dorsiventrally, bearing several sharp 
ridges toward the apex, the walls in median 
cross section 0. 5-0.8 mm. thick, smooth or 
with 1-2 slight ridges, stramineous. 
range: Swampy woods or high bogs at 
3,500 to 4,700 feet altitude and the open bog 
of Alakai Swamp from the Kalalau Pali to 
Keaku Cave; also near Wahiawa Swamp at 
2,100 feet altitude. 
TYPE: Kauai, Waineke Swamp, Kokee, 
June 28, 1926, Otto Degener and Henry Wiehke 
2,187. 
SPECIMENS examined: All from Kauai: 
Kokee, C. Skottsberg 1,020; Kawaikoi Stream, 
C. N. Forbes 908.K; Kalalau Pali, C. N. Forbes 
1,037.K; n. w. end Alakai Swamp, H. St. 
John et al. 10,736; Kauluwehi, J. F. Rock 
12,914; central plateau, trail to Kaluiti stream 
and Keaku cave,/. F. Rock 3,063 (also dupli- 
cates under the temporary numbers 3,075, 
3,077, and 3,078); Kaholuamanoa [ = Kaho- 
luamanu], A. A. Heller 2,806; and near Wa- 
hiawa bog, L. H. MacDankls 6l9> 
The only collector recording the stature of 
the plant was MacDaniels and he gave it as 
"small tree, height 4 meters.” The plant is a 
shrub rather than a tree. 
The diagnostic characters of the two species 
are as follows: 
S. kauaiensis (Degener) St. John has the 
branches often long and wand-like, leafy for 
some distance; axillary hairy tufts 6-7 mm. 
long; peduncle 2-10 mm. long; pedicel 
7-35 mm. long; calyx lobes 5-8 mm. long, 
linear or rarely lance-linear; corolla tube 28- 
35 mm. long; corolla lobes hirsutulous within 
except on hyaline- winged margins; endocarp 
wall in median cross section 0. 5-0.8 mm. 
thick, stramineous, the surface smooth or 
with 1-2 slight ridges. 
S. glabra Hooker & Arnott has the branches, 
or at least the lateral ones, ascending with 
leaves crowded near the stem tips; axil- 
lary hairy tufts 3-5 mm. long; peduncle 
0-3 mm. long; pedicel 13-30 mm. long; 
calyx lobes 2-5 mm. long, narrowly deltoid; 
corolla tube 15-24 mm. long; corolla lobes 
hirsutulous within only for a short distance 
just near the base in the submarginal areas; 
endocarp wall in median cross section 1-2 
mm. thick, dark brown, the surface with 
several prominent sharp ridges. 
Scaevola frutescens (Miller) Krause 
Scaevola frutescens (Miller) Krause is dis- 
tinguished from its varieties by having its 
leaves glabrous from the first or, if sparsely 
puberulent near the midrib below, then gla- 
brate long before maturity. This species is 
common on the outer Leeward Islands of the 
Hawaiian Group, extending from Kure 
(Ocean) Island to Midway, Pearl and Hermes 
to Laysan. Then it occurs on Kauai near Na- 
wiliwili Bay and inland on the slopes of the 
Haupu Range; and it has been found on Ka- 
hoolawe Island. On the shores of the other 
main Hawaiian Islands the plant occurs only 
as the var. sericea, which is abundant and wide- 
spread along the shores. 
Scaevola frutescens (Miller) Krause, var. 
sericea (Forster f.) Merrill 
S. frutescens forma moomomiana Degener & 
Greenwell, ex Degener FI. Haw. 340: 
12/1/49. 
SPECIMENS EXAMINED: Molokai: Kaluakoi, 
Kawakui Road, 600 feet altitude, low bushes 
