Genus Isodendrion — ST. John 
217 
the specimens are in the following: 
BISH = Bernice P. Bishop Museum, Hono- 
lulu. All specimens not otherwise 
assigned are in this collection. 
GH = Gray Herbarium, Cambridge. 
P = Museum National d’Histoire Na- 
turelle, Paris. 
US = United States National Herbarium, 
Washington. 
The maturity or completeness of the speci- 
mens cited is indicated in a following paren- 
thesis by the following abbreviations: 
b. = bud 
11. = flower 
fr. = fruit 
i. = immature 
If. = leaf 
St. = stem 
TAXONOMY 
Family VIOLACEAE 
Genus Isodendrion Gray 
Am. Acad. Arts Sci., Proc. 2: 324-325, 1852; 
U. S. Exploring Exped., Botany, Phanero- 
gamia 15: 92-93, 1854; Atlas pi. 8-9, 1856. 
Shrubs; stems mostly branching, woody, 
the bark roughened by the long persistent in- 
durate subtriangular stipules; leaves alternate, 
short petioled; blades chartaceous to sub- 
coriaceous, from linear lanceolate to elliptic, 
oblanceolate, or ovate, the margins toothed 
towards the apex; inflorescence axillary (a 
reduced cyme), of a single peduncle bibracte- 
olate at apex, bearing a single 1 -flowered 
pedicel; flowers fragrant; sepals 5, unequal, 
persistent, more or less lance-ovate, glabrous 
or pubescent, the margins thin, glabrous or 
ciliate, overlapping at base; corolla clearly 
irregular; petals 5, greenish or whitish, the 
lower two thirds being a ligulate claw, the 
upper third a broader, more or less oval limb, 
the claws laterally imbricate, the uppermost 
exterior, the lower petal interior, the whole 
imitating a corolla tube, particularly the lower 
petal and less so the others have the upper 
half of the claw and at least the base of the 
limb fleshy, thickened, rigid and channel- 
like; petals and stamens marcescent, persist- 
ing around the fruit; stamens 5, unequal or 
subequal, free on the receptacle; filaments 
ligulate, with or without a dorsal enlarged 
nectar gland; anthers oblong to ovate or sub- 
sagittate, the two sacs oblong, dehiscing by 
a longitudinal medial suture; connective in 
all the stamens or at least in the lower ones 
produced into a terminal mucro, flap, or 
prickle, which is usually incurved; ovary 
glabrous, 1 -celled, with 3 placentae bearing 
2-4 orthotropous ovules; style glabrous, fili- 
form, gently sigmoid, the apex decurved and 
enlarged; stigma oblique, deflexed, excavate; 
capsule ovoid, beaked, deeply 3-lobed by 
longitudinal furrows, cartilaginous, promi- 
nently veined, 3-valved; seeds commonly 2 
to each valve, obovate, smooth, shining, the 
apex truncate and excavate; embryo axile; the 
2 cotyledons opposite, oval, free in an ample 
cavity but surrounded by the abundant endo- 
sperm. 
Eourteen species of the Hawaiian Islands. 
KEY TO SPECIES 
A. Sepals glabrous on the midrib; petals glabrous (except in 1. Eorbesii) \ stipules glabrous 
on the midrib or central strip (and in some species on the margin also) ; well- developed 
leaves 5-23.5 cm. long; leaves glabrous, 
B. Sepals glabrous throughout, 
C. Stipules glabrous; blades oblanceolate, 
D. Stipules 2.5 times as long as wide; blades 2. 2-9. 2 cm. long; anthers nearly ob- 
long in outline 6. /. laurifolium 
D. Stipules less than 2 times as long as wide; blades 6.5-22 cm. long, 
E. Lower petal 10 mm. long; stamens not apiculate; leaf margin obscurely 
undulate 1,1. longi folium 
