3 
of Splachnum luteum . 
stance, and this is seen to be more particularly the case in the 
layers of cells next to the foot (Fig. 3 ) 1 . The foot itself is 
seen to consist of a cylindrical mass of parenchyma, with an 
external layer of epidermal cells of a somewhat columnar 
form, which contain a considerable quantity of protoplasm, 
together with large distinct nuclei (Fig. 4). The protoplasm 
of these cells is found to be aggregated towards the peripheral 
surface, the nucleus being usually found in the mass of proto- 
plasm next to the outer wall of the cell. The large vacuoles 
of these cells are traversed by fine protoplasmic filaments. 
These cells, as. well as those of the cortical layer beneath the 
epidermis, contain a number of very small protoplasmic 
bodies, which are found congregated in large numbers round 
the nuclei of the cells, there being also some in other parts 
of the cell, both in the peripheral layer and in the fine proto- 
plasmic filaments traversing the vacuole. In the epidermal 
cells these bodies are particularly numerous, and are found 
principally in the aggregated mass of protoplasm on the outer 
side of the cells. These bodies may, I think, be safely re- 
garded as leucoplastids. From their number and position, I 
am inclined to believe that they are concerned in absorbing 
substances from the tissue of the oophyte for the nourishment 
of the sporophyte. No starch has been found in the foot. 
The general relation of the foot to the vaginula is shown in 
longitudinal section in Fig. 5. 
In the centre of the foot there is a definite central strand 
consisting of two kinds of tissue, an outer phloem-like layer 
of cells containing protoplasm by means of which it is 
probable that organic substance travels, and an inner strand 
of very thin-walled cells without any protoplasmic contents 2 
which conduct the water up the seta (Fig. 6 and 6 bis). In 
the foot the protoplasm of the phloem-like cells is aggregated 
in each cell towards the periphery as in the epidermal cells, 
but there are no plastids present. The strand of thin-walled 
1 [N. B. The cell-contents are not drawn in Fig. 3.] 
2 Cf. Vaizey, loc. cit. The terms leptophloem and leptoxylem have been used 
to indicate these tissues. For fuller explanations, see paper referred to. 
