4 
Vaizey. — On the Sporophyte 
empty cells 1 I have been able to prove in other species of 
Splachnum conveys the water absorbed by the foot up the 
seta into the tissues of the apophysis. 
The seta has a distinct epidermis beneath which there is 
a layer of sclerotic supporting-tissue, and then a layer of 
parenchyma, the two together forming the cortex. In the 
centre is the central strand, which in the lower end of the seta 
has almost the same structure as that described for the cen- 
tral strand of the foot, from which it is distinguished by being 
larger and less distinctly delimited from the cortex (Fig. 7). 
Higher up in the seta there is a large intercellular canal 
formed in the middle of the axile strand of thin-walled empty 
cells which extends for nearly its whole length. This inter- 
cellular space is lysigenous in origin. A similar passage or 
canal occurs in several other species (Fig. 8). 
A longitudinal median section through the umbrella-shaped 
apophysis (Fig. 9, Plate II) shows that the central strand here 
swells out into a large pear-shaped mass of cells which, in the 
mature sporophyte, contain no protoplasm, and even in the 
younger states only a very small quantity with small, incon- 
spicuous nuclei. Chlorophyll-bodies are absent except in the 
two outermost layers of cells, even in the youngest specimens 
observed, and even here there are only a very few. The cells 
are all thin-walled, and cubical in shape, with no intercellular 
spaces between them. In this tissue, which may be regarded 
as a kind of aqueous tissue, large masses of crystalline in- 
organic matter were frequently found. 
Outside the aqueous tissue (Fig. 10) there is a quantity of 
parenchymatous tissue, with numbers of communicating inter- 
cellular spaces. The cells all contain large numbers of 
chlorophyll-bodies. This tissue extends into the umbrella- 
shaped organ. On the upper surface in the proximal region 
the cells are arranged close to one another, and show a dis- 
tinct tendency to an elongation of their axes in a direction 
vertical to the surface, thus forming a palisade-tissue similar 
1 Vaizey : Note on the Transpiration of the Sporophore of the Musci. Annals 
of Botany, vol. I. 
