6 
Vaizey . — (9/2 the Sporophyte 
There are a number of stomata on the upper surface of the 
apophysis, the position of which is shown in the diagram 
(Fig. 9) : one only is drawn in Fig. 10, although a median 
longitudinal section may pass through as many as three. 
Fig. 15 gives a surface view of a stoma from a young apo- 
physis at a time previous to the development of the sur- 
rounding epidermal cells into the radiate arrangement, which 
they assume in the mature apophysis (Fig. 16). The guard- 
cells are quite separate, as in the vascular plants, thus differing 
from the guard-cells of the stomata of the Polytrichaceae and 
Funaria . A transverse section (Fig. 13) through the middle 
of the stoma shows that the form of the lips is quite normal, 
here again differing from the Polytrichaceae 1 (Fig. 18) and 
some other Mosses 2 . The stoma opens into an intercellular 
cavity of considerable size (Fig. 13), the full extent of which 
is seen on comparing longitudinal (Fig. 14) with transverse 
sections (Fig. 13). The guard-cells of the stomata contain 
a considerable number of large and well-defined chloroplastids, 
thus differing from the rest of the epidermal cells. The action 
of Schulze’s solution shows that on the external surface of 
the stomata there is a distinct cuticle (Fig. 17), which extends 
through the stomatal opening and on to the inner surface of 
the guard-cells. The innermost layers of the cell-wall of the 
guard-cells swell up to a much greater extent, and become a 
much deeper blue with Schulze’s solution than do any of 
the rest of the cell-walls throughout the whole of the apo- 
physis. A similar state of things was found in the stomata of 
Polytrichum commune , where cuticularisation has extended 
completely through the whole thickness of the cell-walls 
of the ordinary epidermal cells, but in the guard-cells the 
inner layers swell up with Schulze’s solution, and are 
coloured deep blue. 
A large quantity of starch is formed in the cells of the 
1 Vaizey, on the Anat. and Develop, of the Sporogonium in the Mosses. 
Journ. Linn. Soc. vol. XXIV. 
2 Haberlandt, Beitrage z. Anat. und Phys. d. Lanbmoose ; Jahrb. f. wiss. 
XVII. 1886. [Compare alsoE. Biinger, Bot. Centbl. 1890, Bd. XLII, p. 193, &c.] 
