Puccinia Geranii silvatici , Karst . 33 
This I find is the case with P. Urticae which, also, has not 
a perennial mycelium l . I have not yet succeeded in tracing 
its life-history; but it is certainly not connected with Aecidium 
Urticae , Schum., as I have elsewhere shown. It possibly runs 
a parallel course with P. Geranii silvatici ; but it is more 
probably a heteroecious species. However this may be, I 
have frequently found the teleutospores which are produced in 
autumn (there are no uredospores) germinate freely at that 
time ; and I have recently found that they germinate equally 
after a winter rest. I gathered some leaves bearing tele- 
utospore-pustules in November, and upon floating them on 
water found them freely germinating when scraped off the 
living leaf. Again, on the 19th April following (5 months 
later), and lastly on the 5th July (8 months), spores from the 
same collection germinated very freely. This therefore is 
another excellent illustration of a teleutospore being both a 
Leptopuccinia and a Micropuccinia. 
Before leaving this part of my subject I would draw atten- 
tion to the somewhat tardy setting in of germination in the 
spores even under favourable circumstances. It will have been 
observed that general germination did not set in until the 
spores had been in water about three to four days, indicating 
the essentially halting nature of teleutospores. My usual 
experience has been that if teleutospores are going to germinate 
at all they do so within twenty-four hours, under the conditions 
in which I placed the Geranium-fungus spores. This halting 
germination occurred in all my cultures, and appears to be 
characteristic. Again, it is probable that the teleutospores of 
the first crop, when they exercise their resting properties, can 
do so only for a month or so, while those of the second crop 
must do so for several months. I have unfortunately not been 
able to demonstrate this by actual experiment. There can 
however be little doubt on this point. 
The existence of the fungus in a plant is indicated both by 
the formation of brown spore-pustules on leaves, petioles, and 
1 Journal of Asiatic Society of Bengal, loc. cit., and Scientific Memoirs by 
Medical Officers of the Army of India, Part II, 1887, p. 38. 
D 
