4i5- 
the cells of Euphorbia splendens. 
of proteid granules shown at (c), and (2) the compact spheres 
(a) with strongly marked double contours ; these are highly 
refracting. Possibly (1) is only a stage in the formation of 
(2) . The granules showed Brownian movements and appeared 
to consist of globulin, as they were immediately dissolved with 
10 per cent. NaCl-solution. The more aggregated spheres 
only became clearer in the centre, indicating possibly a 
globulin centre with a more insoluble shell. But the outer 
part also dissolved in dilute acids and was therefore probably 
composed of derived albumin. 
Such are the forms which the proteid contained in this 
species assumes under various conditions. I will next discuss 
their distribution with a view to showing its probable func- 
tion in the economy of the plant. 
Distribution in the stem. This is best seen by a reference 
to Fig. 3, which shows a cross-section of part of the stem. 
The cells containing proteid occur (1) in the inner layers of 
the cortex just outside the bast, (2) in the medullary rays (m), 
(3) in the pith (/) just within the wood. 
The inner parts of the cortex contain large numbers of 
latex-tubes whose thick walls were stained faintly blue when 
treated with Loew’s ferrocyanide-test, a fact which may 
be worth noticing by the way as it tends to support 
Wiesner’s hypothesis of the impregnation of the cell-wall by 
proteid. 
Although the cells surrounding the latex-tubes frequently 
contained proteid, there did not appear to be any such 
definite relation between the two as to suggest that the latex- 
tubes function as channels of distribution for the proteid. 
Scattered irregularly here and there are cells containing 
starch (.$*), which do not, however, form a regular starch- 
sheath. Cells containing proteid do not occur quite so con- 
stantly in the medullary rays and pith as in the inner layers 
of the cortex, but large quantities of proteid were frequently 
found in those parts. 
In quite young stems proteid was frequently observed in 
the peripheral layers of the cortex. 
G g 2 
