in Myzodendron punctulatum, Banks et Sol. 189 
three other vascular bundles, one to each carpel, the carpels 
alternating with the setae. The setae are distinctly hypo- 
gynous and free, but protected at first by the lamellar out- 
growths of the floral wall, as stamens are by the perianth in a 
floral bud. I propose to defer any attempt to assign to the 
setae their morphological value. The ‘ calyx-tube,’ forming 
an epigynous sheath surrounding the base of the style, is, as 
has been said, evascular. On its inner surface are simple 
stomata like those found in the grooves on the outer surface 
of the flower. These stomata are few, occur at the same level 
and at the bottom of slight depressions, and cause the, other- 
wise, entire margin of this c calyx-tube ’ to appear lobed 
slightly. The main floral vascular bundle is very short, stops 
abruptly at the base of the flower (Fig. 3), where it is rather 
the meeting-point of the three carpellary vascular bundles. 
The central strand of tissue of the flower forms a continuous 
tract with the tissue of the base of the placenta. Surround- 
ing this central strand are several layers of thin-walled empty 
cells, the strand itself being formed of thicker walled oblong 
cells (Fig. 4). 
Morphology of the Female Flower. 
As the orientation of the flower is unknown \ I have made 
a floral diagram (Fig. 11). There was no trace of any 
structure which could be regarded as the apex of a lateral 
dwarf branch, on which the two flowers in the axil of the 
scale could arise. The odd seta is posterior, the three carpels, 
ovules, and stylar lobes are opposite one another and alter- 
nate with the setae. The male flower is naked, it may be that 
the female flower is also. The setae are hypogynous and 
arise external to the ‘calyx-tube.’ If the female flower is 
naked the setae may be staminodes (corresponding to the 
rudimentary ovarium in the & flower). Their position 
(orientation |) is, however, different from that in all other 
Santalaceae 2 . If they are staminodes the ‘ calyx-tube ’ 
1 Eichler, Bliithendiagramme, p. 543. 
2 Ibid. p. 537. 
