in Myzodendron punctulatani> Banks et SoL 19 1 
by direct expression), are quite opposed to Van Tieghem’s 
views. The central placenta in Myzodendron is, as apparently 
in other Santalaceae , axial in nature, and carries three ovules. 
Nursing of the Embryo. 
I shall confine my remarks, for the present, to the changes 
in that ovule which is destined to become the single seed of 
the ripe fruit. As a result of fertilisation the oosphere becomes 
surrounded by a cell-wall and converted into the oospore, 
retaining its one-sided position, to the placental side of the 
long axis of the ovule. The secondary nucleus of the 
embryo-sac divides repeatedly to give a row of nuclei extend- 
ing the whole length of the embryo-sac, and soon separated 
from another by cell-walls, so that the interior of the embryo- 
sac is occupied by a uniseriate column of endosperm cells. 
By the time this has happened, and as a result of fertilisation, 
a remarkable change is observable at the narrow antipodal 
end of the embryo-sac, which elongates upwards and back- 
wards in the body of the placenta, then makes a sharp bend 
on itself, and continues its penetrating course, in a more or 
less winding manner, through the free column of the placenta 
and on through the tract of tissue continuous with this until 
it reaches the base of the flower, where its tip dilates and 
becomes imbedded in the vascular cup, formed by the three 
carpellary vascular bundles diverging from the floral vascular 
bundle, between the tip of which and the descended tip of 
the embryo-sac a few layers of rich parenchymatous cells 
intervene (Figs. 12, 13, 14). Throughout the time of its 
prolongation the embryo-sac remains a uniseriate column of 
uninucleate richly protoplasmic cells (Figs. 12, 15), elongating 
apparently by apical growth, intercalary formation of new 
cells being very rare. The tip of the embryo-sac is pointed, 
ensuring a more ready passage through the placental tissue. 
The cross walls, which give rigidity to the penetrating sac 
and tend to prevent its collapse, disappear from the placental 
part of the embryo-sac, except at the dilated (pear-shaped) 
