56 Acton . — On the formation of sugars 
the glands are situated. The fibrovascular tissue is con- 
spicuously developed in the close small-celled septum-paren- 
chyma surrounding the epithem of the glands. 
The cavity is straight and narrow, widest in the central 
portion of the gland, prolonged upwards into the short 
straight duct, and gradually diminishing in width towards 
the lower extremity, till the epithelial cells become almost 
contiguous. 
The epithelial cells are 
somewhat elongated, entirely 
contiguous laterally, but rather 
irregular in shape and size ; 
the free walls bordering on the 
cavity are more or less arched 
outwards, and show no traces 
of cuticle ; they are of the 
same thickness as the lateral 
walls and stain perfectly uni- 
formly with chlorzinc-iodide 
and methylene blue (Fig. 2). 
Several layers of well-marked 
epithem tissue invest the epi- 
thelium on the inner side ; 
those next the epithelium are 
considerably smaller than the 
cells of the septum-parenchy- 
ma ; but the outermost layers 
pass gradually over into the ordinary septum-tissue ; there is 
no endodermis or distinct line of demarcation between 
the epithem of gland and the surrounding parenchyma. 
The epithem-cells are more or less angular and fit closely 
without any intercellular spaces, whereas the surrounding 
tissue is very loose. The connection of the fibrovascular 
bundles with the epithem tissue is very evident ; some of 
the branches run entirely in the epithem, and endings of 
the usual nature can be observed at intervals. Other branches 
run with one side contiguous to the epithem and the others 
Fig. 2. Narcissus Tazetta, L. Trans- 
verse section, showing structure of the 
gland shortly before opening of flower, 
from a section mounted in dilute glycerine, 
and stained with methylene blue, after 
treatment for three hours before staining 
with 10 per cent, aluminium chloride in 
absolute alcohol ; 7 . cavity ; epl. epi- 
thelium ; e. epithem. 
