in the septal glands of Nar cissus. 57 
surrounded by the septum-parenchyma; every variety of 
intermediate stage between bundles which run completely 
in the epithem and completely in the septum-parenchyma 
may be observed. 
It is interesting to note here that it follows from Grass- 
man’s account of the development of septal glands that the 
epithelium is true epidermal tissue, although the epithem is 
derived from the fundamental ground tissue ; and that in 
these structures we find an instance of epidermal cells taking 
on a secretory function also assisted by the adjacent hypo- 
dermal layers, as is so frequently the case in vegetable 
secretory structures. 
Fig. 3. Fig. 4. 
Figs* 3? 4- Appearance of protoplasm in the epithem and epithelium cells of gland 
of Narcissus Tazetta, L. before formation of sugar, from transverse section of a 
young bud. Mounted in glycerine and alcohol after staining with borax- carmine ; 
from material preserved in absolute alcohol. /. position of gland-cavity. 
W. Gardiner 1 compares the structure of nectaries to that 
of chalk-glands occurring in Saxifragaceae and Crassulaceae, 
and this is especially noticeable in the case of septal glands ; 
but it must be remembered that the physiological significance 
of the two cases is quite different, as chalk-glands only 
excrete in virtue of the activity of root-pressure, whereas the 
secretion in nectaries is entirely independent of the same. 
If a transverse section through the upper part of the ovary 
Proc. Camb. Phil. Soc., vol. v, pt. i, and Q. J. Mic. Sci., vol. xxi, N. S. 
