io5 
and Affinities of Trapella. 
agrees in its axile placentation ; but the spines developed on 
the fruit are a non-scrophularineous character. This Order 
gives us but few aquatic members, Limnophila and Hydrotriche , 
and isolated species of larger genera, none of which however 
especially recalls Trapella. 
Bignoniaceae constitute a homogeneous group with their 
winged seeds, and, furthermore, essentially parietal placenta- 
tion, Trapella deviating in both these particulars. 
Trapella has been referred to Pedalineae, since in this Order 
occur forms with few ovules, and curiously-appendaged, lig- 
nified, non-dehiscent fruits. These characters taken together 
with its opposite leaves, solitary axillary flowers, and its 
‘ quaternary vesicles ’ and general disposition, confirm such 
determination. The appendaged fruit forms an especially 
strong analogical character. 
To Myoporineae Trapella approaches very nearly in some 
respects. In the more typical forms of this Order the ovary 
is bilocular, with two pendulous ovules in each loculus 
inserted on the partition. The seeds are often long, with 
only a small amount of endosperm and superior radicle. The 
leaves however are rarely opposite, and the fruit is un- 
appendaged. Our plant must thus be classed with Pedalineae 
rather than here. 
The affinity between Myoporineae and Pedalineae I believe 
may turn out to be a nearer one than that indicated in the 
Genera Plantarum — indeed Lindley (Vegetable Kingdom) 
places them very close together. I shall revert to this matter 
subsequently. 
In view of the heteromorphous nature of Pedalineae, as it at 
present stands, and in view of the possibility of its being 
divided up in the future and its genera relegated to other 
Orders, it may be well to see to what genus, or group of 
genera, now included in Pedalineae, our plant seems most 
nearly allied 1 . 
1 Since the above was written the concluding part of vol. ix. of Baillon’s 
‘ Histoire des Plantes ’ has appeared, containing Scrophulariaceae. Baillon makes 
Pedalineae (with the exception of the Martyneae, which will doubtless be treated 
