205 
on the Saprolegnieae. 
investment surrounding the cell cavity or lumen [often two or 
three vacuoles in Acklya \ ; and (3) starved sporangia, as I 
have elsewhere named them, the ‘inhaltsarme’ of Rothert, with 
only a thin parietal investment of protoplasm and an immense 
vacuole. These differ in the processes of segregation. 
In the full sporange granules gradually wander into the 
hyaloplasma disk which thus becomes indistinguishable. In 
the other forms the granular protoplasm first retracts from 
the disk with which it is only connected by a thin hyaline 
layer investing the wall and a few delicate plasmatic threads, 
so that the lumen is widest at the base of the sporange. 
Vacuoles then appear in the disk, soon enlarging and 
communicating with the main vacuole of the sporange. The 
disk then thins in the centre, and rises peripherally up the 
walls. The granular protoplasm again stretches down towards 
the septum, and finally by the wandering of granules into the 
hyaloplasma the latter loses its character. At the beginning 
of this process, the septum usually bulges towards the basal 
hypha, thus indicating an increase in the turgescence of the 
sporange. At the end of these processes the protoplasm 
usually shows more or less striation or flockiness, due to the 
uneven distribution of granules, and, in unfilled sporangia, 
has an uneven surface towards the lumen. The distribution 
of granules in the protoplasm, and of protoplasm in the 
sporange, gradually becomes uniform. During these stages 
after the formation of the septum the sporange never 
elongates by more than half per cent., irrespective of the 
concavity of the septum and the formation of the ‘process/ 
except in Rothert’s Saprolegnia , sp. 2. 
The ‘ process ’ may be formed even before the septum, at 
the same time with the differentiation of the spore-origins, or 
most frequently between these two formations. It usually 
occupies the apex of the sporange, but may develop at any 
point [except the septum]. Here again hyaloplasma ac- 
cumulates at a spot, bulging out the membrane ; the bulging 
of the membrane continues with the accumulation of hyalo- 
plasma, until a short cylinder with a nearly hemispherical top 
