310 Hartog . — On the Floral Organogeny 
imbricate quincuncially (sep. i antr.). The four sepals become 
free at the rim of the long floral tube, on which are also 
inserted the five (variably) imbricate petals, not markedly 
heteromorphic or unequal, and nine or eleven 1 stamens mona- 
delphous, the long erect tube split to the base on the posterior 
(vexillary) side. The solitary carpel has the normal orien- 
tation of the Leguminosae, its dorsal suture being anterior, 
its placenta posterior ; its stipe is adnate as a ridge to the 
posterior edge of the floral tube, and becomes free at the 
posterior edge just within the vexillary petal. 
Saraca has the calyx, gynaeceum, and floral tube of Brownea ; 
but it is free, only slightly equitant. The flower is always 
apetalous, and has only the seven anterior stamens, which are 
free to their base ; an eighth sometimes occurs, it is then pos- 
terior. The inflorescence is a stiff panicle twice or thrice 
branched, from the old wood. The bracts of the base of the 
main peduncle are distichous, but higher up they are spiral 
with the divergence § as they are (with antidromy) on the 
secondary and tertiary axes. The upper bracts of the primary 
and secondary branches and all the tertiary are axillant to 
flowers. 
I. The Floral Development of Brownea and 
Saraca. 
The young inflorescences of Brownea , like so many closely- 
packed racemes, contain flowers nearly of the same age. These 
appear first in the axils of the lowest of the spiral bracts, 
and follow in rapid basifugal succession, the main axis often 
bearing finally a terminal flower. The lower floral bracts 
occasionally bear flowers, which, however, take origin much 
later than those in the axils of the spiral bracts. The bract- 
lets appear right and left in quick succession, one a little 
before the other. They soon become connate, i. e. their bases 
are confluent and rise up so as to form a bag of truncated 
1 Ten is a very rare number to find ; eleven is commoner in B . coccinea, nine in 
B. grandiceps. 
