and A natomy of Brownea and Saraca. 3 1 1 
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oc 
conical form, with only a median slit at the apex, the margin 
of the older (Fig. 14, a) bractlet overlapping the other (Fig. 
14, 0). Within this sac the receptacle has widened and 
become obconical, somewhat tilted outward, so that the 
rounded upper surface, covered by the slit of the bracteolar 
sac, looks towards the hollow of the bract ; hence in Brownea 
we have not the same marked difference of pressure between 
the anterior and posterior sides of the flower that usually 
prevails at this stage in closely-packed inflorescences. The 
sepals appear in quincuncial order ; sepal 1 is anterior (see 
Fig. 34), 2 postero-lateral on the 
side of bractlet a, 3 antero-lateral on 
the (3 bractlet side, 4 antero-lateral 
on the a side, 5 postero-lateral on 
the 3 side again ; 2 and 5 early be- 
come confluent at the base ; but the 
resulting posterior member is long, 
unequally divided by a notch into 
a larger portion corresponding to 2, 
and a smaller to 5. 
The petals arise as a simultaneous 
whorl, and so do the alternipetalous 
outer stamens. At this stage the 
swollen centre of the floral re- 
ceptacle has become excentric to the flower, coming close up 
to the base of the vexillary petal behind. Hence we usually 
find in B. grandiceps only four antipetalous inner stamens, 
there being no room left for a posterior stamen before the 
vexillum ; in B. coccinea , however, two are formed here ; we may 
regard this as a true case of chorisis due to the pressure of the 
gynaeceal tubercle. This now rises up as a horseshoe with 
its limbs closely approximated behind. The staminal sheath 
forms late, and so do the receptacular tube and gynophore. 
In Saraca the bractlets soon overlap above the young 
receptacle, on the side of the bract, but leave a small gap 
at the posterior side of the flower until the appearance of the 
sepals ; they never become connate, though a overlaps /3 on 
Fig. 14. Floral diagram of 
Brownea coccinea. Saraca has the 
same diagram with the omission 
of the petals and the four posterior 
stamens. 
