312 Hartog. — On the Floral Organogeny 
the posterior as well as anterior side of the flower. The 
receptacle is nearly erect, not tilted outwards. The calyx 
develops as in Brownea , but the confluence of the sepals, 
2 and 5 , is earlier and more complete. I have failed to see 
any sign of petaline tubercles. The stamens appear in 
ascending order in an antero-posterior direction in rapid 
succession. First, an anterior stamen, then the pair next to 
it, then another pair, and then another; between these, on 
the posterior side of the flower, is seen a narrow ridge which 
soon shows three minute tubercles ; the middle one may 
enlarge somewhat, but their identity and presence are soon 
lost to view. At the time that there are five stamens, the 
central tubercle shows a slight annular margin which almost 
immediately becomes horseshoe-shaped owing to uneven 
growth ; this is the carpel. 
Now the important point in the above developments is, fhat 
in two closely allied genera the flower of the one develops 
as one would say ‘ normally,’ by successive whorls, and in 
the other genus there is a marked acceleration of the anterior 
side. Considering the flowers of these two genera alone, it 
might be thought that Saraca , the more modified form of the 
two, with its apetalous flowers and reduced androecium, owed 
to these points a distorted ontogeny. This conclusion, how- 
ever, is unsustainable. In all Leguminosae hitherto studied, 
including the actinomorphous Mimoseae, the flower is known 
to develop from below upwards, with the appearance of the 
parts next the bract accelerated, those next the inflorescence- 
axis retarded. In this order then Saraca is normal, Brownea 
is exceptional ; and it is the eucyclic development of Browjiea 
which calls for explanation. We must, therefore, regard this 
eucyclic development as secondary, and hold that a reversion 
has taken place here, conditioned by the nearly actinomorphous 
relations of the adult flower. With this is correlated the 
peculiar position of the flower-axis (at right angles to the 
inflorescence-axis) at the stage when the flower-leaves make 
their first appearance. 
It is obvious that the flowers of most Leguminosae must 
