and Anatomy of Brownea and Saraca. 315 
of the tube, which is now visible. A little higher up the ring 
at the posterior side of the flower gives off three bundles : one 
posterior for the (absent) vexillum, two lateral to the two 
posterior staminal rudiments (mostly undeveloped) in the adult 
flower ; the two latter are inconstant. The lateral gaps soon 
close up, but the posterior is continued upwards as the gap of 
the ventral suture of the carpel. To state the case in other 
words : — the leaf-trace of the carpel forms a horseshoe with the 
opening posterior. The trace of the (absent) vexillum closes 
this gap, and the two posterior staminal traces enter the sides 
of the cylinder thus formed. Lower down the cylinder opens 
out on its anterior side ; and the posterior arc so formed 
receives in its flanks the insertion of the traces from the 
posterior sepals (2 and 5). The traces of the remaining nine 
stamens are concentric above ; below each opens into two, and 
between the two of each stamen is inserted the trace of one 
of the remaining sepals or petals. In the upper part of the 
tube the leaf-traces of the sepals and (absent) petals branch 
collaterally and approximate irregularly, so that each leaf 
receives a number of bundles. 
In Brownea the arrangement of the bundles is also a nine- 
pointed festoon with the odd bay posterior. The points go to 
the nine anterior flower-leaves, i. e. to the sepals and the four 
anterior petals. The flanks of each of the seven anterior cusps 
bend inwards, and converge in pairs to the seven anterior sta- 
mens. The remaining small bundles, one from each of the six 
anterior bays, cross obliquely towards the back of the flower, 
and form a crescent open behind for the posterior part of the 
gynophore ; the flanks of the two anterior cusps also converge 
to form each a posterior staminal trace, smaller than the seven 
anterior. The posterior bay now becomes convex behind, and 
with the large bundles from the two adjacent bays and the 
crescent formed of the six small bundles from the lateral and 
anterior bays, forms a circle at the posterior side of the flower, 
separated from the excentric arc of the seven more anterior 
staminal traces by the opening of the tube. Higher up the 
vexillary trace (flanked by two small crescents for the tenth 
