3 1 6 Hartog . — On the Floral Organogeny 
and eleventh stamens in B. coccinea) separates from the rest of 
the circle which is continued in the gynophore. In other 
words, each staminal trace splits into two £ half-traces/ which 
are inserted on either side of the corresponding sepaline or 
petaline trace ; with the exception that when eleven stamens 
are present, the traces of the posterior pair go without split- 
ting to either side of the vexillary trace. The traces from 
the gynophore are inserted between the £ half-traces ’ of ad- 
jacent stamens, from this point of view the two posterior of 
eleven stamens behave as £ half-traces ’ ; the anterior traces 
from the gynophore are smallest. 
Further, we must note that irregularities often occur ; espe- 
cially that the small traces of the gynophore from the seven 
anterior stamens may be much reduced, and I think in some 
cases absent. 
Let us see what morphological light we can get from this 
study. First of all, Saraca, though apetalous, is equipped with 
a full set of alternisepalous traces obviously equivalent to the 
petaline traces of Brownea. Here we have evidence of the 
phylogenetic abortion of the petals, such as could not have 
been gleaned from the ontogeny. Again the double nature of 
the posterior sepal in both genera is confirmed. Then the 
congenital chorisis of the two posterior stamens (completing 
eleven) in Brownea coccinea , inferred from comparison with other 
Leguminosae, is confirmed by the fact that either sends down 
a single trace to be inserted on one flank of the vexillary 
trace ; the others send down a trace which divides into two 
£ half-traces/ going to either flank of a floral leaf-trace. 
All this is plain sailing ; but it is otherwise when we look at 
the gynophoral traces of Saraca , taken by itself. 
Here the stipe of the pistil receives its traces only from the 
posterior side of the vascular cylinder of the pedicel ; in 
other words, they enter between those of the other floral 
organs of the posterior side only. If we consider the stipe 
as an internode between the stamens and pistil, it is obvious 
that its components should be inserted symmetrically between 
those of the lower verticels. If, on the contrary, with Karl 
