and A natomy of Brownea and Saraca. 3 1 7 
Schumann, we regard it as the petiolar base of the carpellary 
leaf, since the carpel is certainly anterior, its bundles should 
join those of the anterior organs. Taken alone, the distribution 
of the leaf-trace bundles of Saraca , instead of shedding light 
on the morphology of the gynophore, presents an enigma for 
solution. 
In Brownea and the Proteaceae we find an explanation of 
the enigmatical conditions of Saraca . The proteaceous flower 
has a four-leaved perianth with antiphyllous stamens ; and 
a single stipitate carpel with its placenta posterior — conse- 
quently the carpel itself is anterior, as in Saraca , Brownea , and 
the other Amherstieae. In some species the flower is actino- 
morphic ; in others the perianth is open to the base in front, 
gamophyllous and gibbous towards the back, so as to form 
a short tube on the side next the ventral suture of the carpel, 
not the dorsal as in Amherstieae. In the actinomorphous 
species I have examined the leaf-traces of the gynophore are in- 
serted symmetrically ; in the zygomorphous we may distinguish 
two cases. In some ( Grevillea spp.) the leaf-traces from the 
posterior side of the flower are present, but weaker ; in the other 
case ( Stenocarpus salignus ) they are absent, and the bundles 
are exclusively derived from the anterior side of the flower. 
A judgment founded exclusively on the flower-anatomy of 
such a flower would say that the carpel being anterior received 
its bundles from the anterior side of the flower ; but the case 
of Saraca , where the stipe of the anterior carpel receives all its 
traces from the posterior side, prevents this easy solution. 
But the true conditions are obvious ; the excentric position 
of the stipe, owing to the formation of the flower-tube, has in- 
fluenced the internal anatomy. With the tube posterior, the 
posterior bundles are reduced or absent, as in Proteaceae 1 ; 
with the tube anterior, the anterior leaf-traces are reduced 
( Brownea ) or absent (Saraca). 
1 From examination of some specimens of a Bauhinia kindly sent by Prof. 
Oliver, I believe the conditions are the same as in Stenocarpus salignus , the tube 
being anterior to the adnate stipe ; but I failed to obtain satisfactory prepara- 
tions from the dry material. 
