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PACIFIC SCIENCE, Vol. XIV, April I960 
Fig. 7. Dolioletta valdiviae, stomach, intestine and 
ventral process of phorozooid. 
For the specific distinctions to be maintained, 
reliance is being placed on differences of pro- 
portional widths of muscle bands. It is probable 
that the amphiclinous condition with the myo- 
plane ( Garstang’s terminology) between M.3 
and M.4 would persist among other species of 
a particular subgenus (in this instance Dolio- 
letta ); and, within the condition imposed by 
this, it is probable also that species may be dis- 
tinguishable by differences in the proportional 
widths of muscle bands, much as different rela- 
tive widths of muscles, combined with a change 
in position of the myoplane, occurs between 
Dolioletta and Doliolina. 
The occurrence together of "old nurses,” 
gonozooids, and phorozooids does not necessar- 
ily constitute evidence of identity. However, the 
"old nurses” from New Zealand consistently 
differ from those of a second species in the sub- 
genus, and it is reasonable to suppose in the 
absence of other Doliolids, that the nurses, 
gonozooids, and phorozooids belong together. It 
is proposed that they be considered as those of 
Dolioletta valdiviae. 
Garstang has been concerned that specimens 
from the area of the Benguela Current which he 
called D. valdiviae might be traceable to "Some 
'abnormality’ of growth conditions . . .” and that 
"the whole of the valdiviae stock may have been 
the produce of a single swarm of nurses reared 
under peculiar conditions of temperature and 
food supply.” With the finding of specimens 
from New Zealand similar to those Garstang 
discusses, this possibility is reduced. That closely 
similar conditions of temperature and food sup- 
ply prevailed in the two regions, and that re- 
actions to these of the produce of some old 
nurses were the same, while not impossible, is 
unlikely. 
Order DESMOMY ARIA 
Family SALPIDAE 
Genus Iasis (Lahille) 
Iasis zonaria (Pallas) 
Salpa nitida Herdman, 1888 
Salpa zonaria cordiformis Ritter, 1905 
Salpa zonaria Metcalf, 1918; Thompson, 1948; 
Sewell, 1953 
Fourteen specimens were collected from five 
stations, between Cook Strait (January) and 
Station 826, in the subantarctic. 
Metcalf (1918) and Thompson (1948) de- 
scribe and discuss this species at length. Figures 
9 and 10 indicate the chief diagnostic features 
of the solitary and aggregate forms. As well, in 
solitary forms, the test is rigid, the muscles are 
broad and interrupted ventrally, and the body is 
symmetrical. In aggregate forms, the test is 
rigid, the muscles are widely interrupted ven- 
trally, and the body is asymmetrical. Often one 
or more embryos are contained within the body. 
The largest specimen was 63 mm. long, the 
smallest, 29 mm. 
Genus Thalia Blumenbach 
Thalia democratica Forskal 
Thalia democratica mucronata Herdman, 1888 
FIG. 8. The proposed "old nurse” of Dolioletta val- 
diviae, a 29 mm. specimen. 
