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PACIFIC SCIENCE, Vol. XIV, April I960 
16(15). Rostrum reaching to end of anten- 
nular peduncle 
.A. lamellifer Kubo 
Rostrum at most reaching to end of 
second antennular article, usually 
shorter 17 
17(16). Palm of chelae 2.5 times length of 
carpus, or shorter, in mature 
females A. marshallensis Chace 
Palm of chelae 3 to 5 times length 
of carpus in mature females 
A . rhothionastes Banner 
18(15). Infracorneal tooth longer than ex- 
tracorneal tooth 19 
Infracorneal tooth shorter than ex- 
tracorneal tooth 20 
19(18). Dorsal carina extending along an- 
terior half of carapace and ros- 
trum A. japonicus Kubo 
Anterior carapace without dorsal 
carina . A. esakii Kubo 
20(18). Dactylus of large chela of male 
Strongly curved, "half-moon 
shaped," much longer than fixed 
finger..... A. oshimae Yokoya 
Dactylus of large chela of male at 
most slightly curved, not "half- 
moon shaped," not markedly 
longer than fixed finger (males 
unknown in A. haswelli below) ..21 
21(20). Carpus and chelae slender in fe- 
males 5 A. haswelli Coutiere 
Carpus and chelae of females more 
robust 5 .... A. minikoensis Coutiere 
22(1). With well-developed supracorneal 
teeth 23 
Without trace of supracorneal 
teeth; carpus of second legs with 
four articles 25 
23(22). With carpus of second legs with 
five articles, lower portion of 
palm of large chela bearing nu- 
merous papillae 
A. verrucosus Banner 
5 Co”tiere’s only differentiation between these two 
species is as follows: "[The cheliped of A. hasivelli ] 
a la mem; forme cylindrique et grele [as A. dimor- 
phus\ ce qui distingue l’espece de V A. minikoensis ou 
la oirce est beancono plus robust et le carpe court.” 
Bull. Soc. Philom. IX, 11(5): 2. 1908. 
With carpus of second legs with 
four articles; lower portion of 
palm of large chela without 
papillae 24 
24(23). Ventral side of rostrum smoothly 
convex; base of outer flagellum 
with four articles; propodal spine 
of uropod simple 
A. borradailei (Coutiere) 
Ventral side of rostrum with an ob- 
tuse angle; base of outer anten- 
nular flagellum with more than 
four articles; propodal spine of 
uropod bifid 
A. ghardaqensis (Ramadan) 
25(22). Rostrum reaching to middle or end 
of third article of antennular 
peduncle. Usually associated with 
sea urchins of the genus Echi- 
nometra A. indicus (Coutiere) 
Rostrum not reaching past middle 
of second antennular article, 
usually only to end of first article 
or shorter 26 
26(25). Telson elongate, 6.5 times as long as 
posterior margin is broad; pos- 
terior margin one-third breadth 
of base. Symbiotic association, if 
any, unrecorded 
A. amboinensis (de Man) 
Telson less than 4 times as long as 
posterior margin is broad; pos- 
terior margin one-fourth to one- 
fifth breadth of base. Usually as- 
sociated with sea urchins of the 
genus Heterocentrotus 
A. dorsalis (Stimpson) 
Athanas areteformis Coutiere 
At h anas naifaroensis Coutiere, 1903- Bull. Soc. 
Philom. Paris IX, 5(2): 77, figs. 14-16; 
[same description and figures in 1905, Fauna 
and Geog. Maidive and Laccadive Archi- 
pelagoes 2(4): 859, fig. 131 a-c]. 
Athanas areteformis Coutiere, 1903. Op. cit. p. 
79, figs. 17-19; [same description and figures 
in 1905, op. cit., p. 860, fig. 132 a-c], 
Athanas erythraeus Ramadan, 1936. Bull. Fac. 
Sci. Egypt. Univ. 6: 13, pi. 1, fig. 1. 
