Alpheid Shrimp, Y— Banner and Banner 
145 
TABLE 3 
Variation in the Chelipeds of Athanas rhothionastes sp. nov. 
LOCALITY 
SEX 
CARA- 
PACE 
LENGTH 
SMALL CHELA 
LARGE CHELA 
Fingers ^ 
g V | 
3 c 1 
proportion to 
palm) 
Palm 1/b 
Carpus 1/b 
Merus 1/b 
J3I 
3’ 
DTP 
n> 
in 
v> 
(in 
$ O g 
3 2 
£ 
proportion to 
palm) 
Palm 1/b 
Carpus 1/b 
Merus 1/b 
Tahiti 
$ 
2.8 mm. 
0.5 
1.0 
1.6 
1.4 
2.7 
2.5 
3.2 
0.4 
1.0 
0.2 
0.8 
2.7 
1.2 
2.8 
Tahiti 
9 
3.2 mm. 
0.3 
1.0 
0.2 
1.0 
2.6 
1.3 
2.5 
0.4 
1.0 
0.2 
0.8 
2.7 
1.2 
2.5 
Tahiti 
9 
3.4 mm. 
_ 
— 
— 
— 
— 
— 
— 
0.4 
1.0 
0.2 
1.0 
2.8 
1.5 
2.8 
Canton 
9 
3.8 mm. 
0.3 
1.0 
0.2 
0.8 
2.9 
1.2 
2.8 
0.4 
1.0 
0.2 
1.0 
3.1 
1.4 
3.2 
Canton 
9 
3.9 mm. 
0.3 
1.0 
0.2 
0.8 
3.0 
1.3 
3.0 
0.3 
1.0 
0.2 
0.9 
2.8 
1.1 
3.6 
Eniwetok 
9 
4.1 mm. 
0.4 
1.0 
0.2 
0.9 
2.7 
1.4 
2.9 
0.3 
1.0 
0.2 
0.9 
2.6 
1.5 
2.7 
Canton 
9 
4.3 mm. 
0.4 
L0 
0.2 
1.0 
2.7 
1.3 
2.7 
— 
— 
— • 
— . 
— 
— 
— 
Canton 
9 
4.5 mm. 
0.4 
1.0 
0.3 
1.1 
2.6 
1.8 
2.8 
_ 
— ■ 
_ 
— 
— - 
— 
— 
Samoa 
& 
2.7 mm. 
— 
■ — 
— 
_ 
— 
— 
— 
0.4 
1.0 
0.3 
1.1 
2.8 
1.8 
3.2 
Tahiti 
S 
2.8 mm. 
0.3 
1.0 
0.3 
1.0 
3.0 
1.5 
2.7 
0.4 
1.0 
0.2 
0.9 
2.5 
1.0 
2.5 
Fiji 
$ 
3.2 mm. 
— 
— 
— 
— 
— - 
— 
— 
0.2 
1.0 
0.2 
0.9 
3.0 
1.6 
2.4 
Tonga 
$ 
3.4 mm. 
0.3 
1.0 
0.2 
0.9 
2.9 
1.6 
2.6 
0.3 
1.0 
0.2 
0.9 
2.6 
1.6 
2.9 
Canton 
s 
3.4 mm. 
0.4 
1.0 
0.2 
1.0 
2.8 
1.5 
3.3 
0.3 
1.0 
0.2 
0.9 
2.9 
1.3 
2.5 
Tahiti 
s 
3.5 mm. 
0.3 
1.0 
0.1 
0.9 
3.2 
1.6 
3.0 
— 
— 
— 
— 
— 
— 
— - 
Tahiti 
s 
3.7 mm. 
0.3 
1.0 
0.2 
1.1 
2.7 
1.0 
2.9 
0.3 
1.0 
0.2 
1.0 
3.1 
1.2 
2.8 
Canton 
$ 
4.2 mm. 
— 
— 
— 
— - 
— 
— 
— 
0.2 
1.0 
0.2 
0.9 
3.1 
1.0 
2.1 
Canton 
6 
4.3 mm. 
0.3 
1.0 
0.2 
1.0 
3.0 
1.5 
2.4 
0.3 
1.0 
0.2 
0.9 
2.5 
0.9 
2.4 
opposition to large tooth (evidently these are 
thin and used for cutting as they slide past each 
other). 
Carpal articles of second leg with ratio: 10: 
1: 1: 1: 5. 
Ischium of third legs unarmed, 0.4 as long as 
merus; merus unarmed, 5 times as long as broad; 
superior distal margin of carpus projecting but 
rounded, carpus 0.5 as long as merus; propodus 
0.9 as long as merus, armed with scattered spines 
along inferior margin; dactylus slender, slightly 
curved, acute and simple, and 0.3 as long as 
propodus. 
Telson 33 times as long as broad distally, 
twice as broad proximally as distally; lateral 
margins with uniform taper; tip broadly arcuate; 
dorsal and terminal spinules normal. Uropods 
longer than telson, outer uropod armed with 
tooth and movable spine at articulation. 
Color in life not noted. 
DISCUSSION : Variation in this series of speci- 
mens has been carefully studied to determine if 
they are distinct from closely related species. 
The rostrum is always acute, almost always ta- 
pering, although there are a few with straight 
parallel sides in the proximal third to half; the 
tip reaches from the middle to the end of the 
second antennular article except for a very small 
specimen from Samoa, where the tip reaches to 
the middle of the third antennular article. In 
many, the rostrum appears more narrow than in 
the holotype and often has a slight upturn 
towards the tip when seen in side view. Like- 
wise, with few exceptions, the extra- and infra- 
corneal projections are quite uniform; in the 
small specimens from Samoa assigned to this 
species the infracorneal tooth is almost acute; in 
one from the type locality the extracorneal tooth 
is bifid on one side and with the infracorneal 
projection rounded but on the other side in the 
same specimen the upper tooth is normal and 
the infracorneal tooth is acute. 
Fig. 3. Athanas rhothionastes Banner. Holotype (female) a-i; allotype j-m. a, b, Anterior region; c, d, e, f, 
chelipeds; f, interior lateral surface of chelae; g, second leg; h, third leg; i, telson; j, k, large cheliped and 
fingers; l, m, small cheliped and fingers; n, large and small chelae of 2.7 mm. (carapace length) ovigerous 
female from Tahiti (BD-2) ; o, same from 3.0 mm. ovigerous female from Eniwetok (BE-4). (Anterior body 
region and telson, scale a; all other appendages scale b.) 
