Alpheid Shrimp, Y— Banner and Banner 
veloped," with only three rows of light setae 
found distally. 
Abdomen normal except anterior pleura 
greatly expanded, posterior pleura small; pleura 
of sixth segment articulated. Telson normal, 
sides almost straight, tapering, 2.2 times as 
broad anteriorly as posteriorly, 4 times as long 
as broad posteriorly; tip extended and rounded. 
Uropods normal, narrow and long. 
Branchial formula including five pleuro- 
branchs, no arthrobranchs, and mastigobranchs 
extending back to second thoracic leg, seto- 
branchs to third. No anal tubercles. 
DISCUSSION: In the old genus Athanas this 
species is closely related to none; in the old 
genus Arete, now united with Athanas, this 
species is most closely related to A. borradailei 
(Coutiere, 1903: 9) and A. ghardaqensis (Rama- 
dan, 1936: 15). From both of these species it 
is separated by the five- jointed carpus and the 
presence of papillae on the palm. The difference 
in the chelae could be from growth or sexual 
dimorphism. A point for point comparison re- 
veals other more minor differences. This species 
differs from A. ghardaqensis by a greater num- 
ber of articulations in the base of the inner 
flagellum, by the presence of a small tooth on 
the basicerite, and by longer uropods. From 
A. borradailei, this species is distinguished by: 
1. A more sinuate lower margin of the rostrum 
and a triangular shape (when seen in dorsal 
view); 2. the presence of serrations on the dis- 
tal ends of the antennular articles; 3. the pres- 
ence of many articulations in the antennular 
flagellum; 4. slight differences in the last article 
of the external maxilliped; 5. on the chelipeds, 
the presence of spines on the ischium, and slight 
differences in the proportions of the merus and 
carpus in addition to the presence of papillae 
on the palm; 6. a broader merus of the third 
legs, being less than 2.5 times as long as broad 
instead of three times. Also A. borradailei is 
reported to lack all thoracic epipodites, while 
this species has masigobranchs on the base of 
the chela and second legs. Neither Coutiere nor 
Ramadan discussed the mouthparts. 
Each of these three species were described on 
the basis of a single individual. While the dif- 
ferences listed above, especially in the carpus of 
the second legs, the chelae and the branchial 
149 
formulae may be valid for their separation, the 
three specimens are so similar in general con- 
figuration and in the structure of the appendages 
and rostral front that their specific separation is 
very doubtful, particularly in view of the varia- 
tion in other species of this genus. However, 
with only one specimen of each nominal species, 
it appears to be safest to leave them as separate 
species. 
The name verrucosus refers to the "warty” 
condition of the palm of the large chela, a char- 
acter in which this species appears to differ from 
both related species. 
Athanas indicus (Coutiere) 
Arete dorsalis var. indicus Coutiere, 1903. Bull. 
Soc. Philom. Paris IX, 5(2): 85. 
Arete indicus Coutiere, 1905$. Fauna and Geog. 
Maidive and Laccadive Archipelagoes 2(4): 
863, figs. 134-135. 
Arete iphianassa de Man, 1910a. Tijdschr. Ned. 
Dierk. Ver. II, 11(4): 312; 1911, Siboga 
Exped. 39a 1 (2) : 164, pis. 3, 4, fig. 11; 1922, 
Siboga Exped. 39a 4 (5): 22, pi. 3, fig. 11. 
Banner, 1956. Pacif. Sci. 10(3) : 325. 
Arete intermedins Yu, 1931. Bull. Mus. Hist. 
Nat., Paris II, 3(6): 513-514, fig. 1. 
Arete kominatoensis Kubo, 1942. Dobutsugaku 
Zasshi fZool. Mag., Tokyo] 54: 82-85, figs. 
1 , 2 . 
DISCUSSION: The habitat of 15 of 16 speci- 
mens from Aitutake was carefully observed 
while the collections were made. They came 
from the inner, but growing, portion of a wind- 
ward reef, in a pool about six feet deep; every 
specimen was associated with the boring sea 
urchin, Echinometra, and were found both on 
the body of the urchin between the spines and 
within the smooth rounded cavity produced by 
the urchin. While most of the other specimens 
were not as carefully observed during their col- 
lection, the ones that were noted (as from 
Eniwetok) were so associated. Evidently, not 
all individual specimens of Echinometra have 
shrimp in association, for Echinometra was fre- 
quently encountered while collecting and the 
shrimp are quite uncommon in the collection. 
For the following discussion of variation 
within the species, it was considered best to 
