228 
PACIFIC SCIENCE, Voi. XIV, July I960 
describes the distinctive stigma of this species 
and section. Also in Martelli’s diagnosis (p. 369) 
he used the words, "stigma . . . pulvinatum.” 
Consequently, the emendation in spelling made 
by Kanehira seems permissible under the Inter- 
national Code of Botanical Nomenclature. The 
section contains two species from Borneo. 
Rykia (De Vriese) Kurz, Jour. Bot. Brit, and 
For. 5: 101-104, 1867 (as Ryckia ); Asiat. Soc. 
Bengal, Jour. 38(2): 147, 1869. Genus Rykia 
De Vriese, Akad. Amsterdam, Verh. 2: 203, 
1854, and in Hooker’s Kew Jour. 6: 268, 1854. 
The alteration to Ryckia De Vriese emend. Balf. 
f., Linn. Soc. Bot., Jour. 17: 35, 48, 1878, and 
Martelli, Webbia 4(1): 96, 1913, was un- 
justified. As De Vriese stated, the genus was 
named in honor of Vice-Admiral J. C. Ryk. The 
Index Kewensis has already judged this change 
in spelling by Balfour as a mistake. As a new 
synonym, is included the section Epiphytici 
Martelli, Webbia 4(1): 96, 1913; Soc. Bot. Ital., 
Bui. 304, 1904. Its single species, P. epiphyticus 
Martelli, from Borneo, has all the morphological 
characteristics of the section Rykia. The fact that 
it was found growing perched upon another 
plant, is considered of no taxonomic significance. 
Type: P. furcatus Roxb. The section contains 
74 species and ranges from Madagascar to Aus- 
tralia: Madagascar (3 species), India (13), 
Ceylon ( 2 ) , Burma ( 5 ) , Thailand ( 3 ) , Malaya 
(9), Indochina (9), China (2), Philippines 
(5), East Indies (22), New Guinea (1), and 
Australia ( 1 ) . 
Rykiella Pic.-Ser., Inst. Sci. Madagascar, Mem. 
B, 3: 155-158, 1951. Type: P. macrophyllus 
Martelli. This and the one other species occur 
in Madagascar. 
subgenus Pandanus 
The use of the name Pandanus for this sub- 
genus is required by the International Code of 
Botanical Nomenclature, since it contains the 
type section and the type species of the genus. 
It has long been known by the sectional name 
Keura (Forsk.) Kurz, Jour. Bot. Brit, and For. 
5: 105, 1867. Type: section Pandanus. Drupes 
or phalanges all or nearly all several-celled. 
Acanthostyla Martelli, Soc. Tosc. Sci. Nat., 
Atti, Proc. Verb. 42: 57, 1933; Inst. Sci. Mada- 
gascar, Mem. ser. B, 3: 11, 122-126, 1951. 
Type: P. spinifer Warb. The section contains 
13 species of Madagascar. 
Australibrassia, sect. nov. Phalangiis pluri- 
carpellosis lobatis in apice valde angustatis, api- 
cibus carpellorum anguste deltoideis vel conicis 
exterioribus inflexis et valde asymmetricis. 
Phalanges much narrowed towards the apex, 
and there lobed between the numerous carpels; 
apices of the carpels narrowly deltoid or nar- 
rowly conic, the outer ones inflexed and very 
asymmetric. 
HOLOTYPUS: P. conicus St. John. Its descrip- 
tion follows. Also in this section is the new 
P. cochleatus St. John. 
Coronata Martelli, Univ. Calif. Publ. Bot. 
12(10): 359, 1930. Lectotype: P. corallinus 
Martelli, 1. c. 395, pi 45, fig. 10-12, 1930. The 
lectotype is from Tongatabu, Tonga. The other 
species, P. W hitmeeanus Martelli, is known from 
Tongatabu, and from Savaii, Samoa; and P. escu- 
lentus Martelli is from Luzon in the Philippines. 
Dauphinensia, sect. nov. Carpellis ( 1 ) - 
pluribus, phalangiis integris in apice late con- 
vexis, stigmatibus omnibus (vel omnibus unico 
centrale 'excepto) in margine circulo vel poly- 
gono apicale. 
Carpels ( 1 ) - several; phalanges entire and 
with the apex wide convex; stigmas all (or all 
except the single central one) on the margin of 
an apical circle or polygon. 
HOLOTYPUS: P. dauphinensis Martelli in Mar- 
telli and Pichi-Sermolli, Inst. Sci. Madagascar, 
Mem., ser. B, 3(1): 38-40, fig. 5, 1951. 
The section contains six species, including: 
from east Africa, P. Stuhlmanni; from the Sey- 
chelles, P. Hornei; from Madagascar, the type, 
and P. concretus, and P. reflexus; and from 
Guam, P. Kafu. 
Hombronia (Gaud.) Warb., in Engler’s Pflan- 
zenreich IV, 9: 50, 1900; genus Hombronia 
Gaud., Bot. Voy. La Bonite, Atlas, t. 27, fig. 17, 
(1843) [= 1841]. The name was published by 
Gaudichaud without a description, but the genus 
was monotypic, and it was illustrated by a figure 
with details, so the name was valid. Type: Hom- 
bronia edulis Gaud., but the epithet is pre- 
occupied in Pandanus , so the valid name is now 
P. dubius Spreng. The section contains 22 spe- 
cies, and it occurs from the Andaman Is. to 
