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Samoa, as follows: Andaman Is. (1 species), 
Nicobar Is. ( 1 ) , Malaya ( 1 ) , Thailand ( 1 ) , 
Philippines (2), Micronesia (3), Indonesia 
(1) , New Guinea (8), Solomon Is. (2), New 
Caledonia (8), Isle of Pines (1), Loyalty Is. 
(2) , New Hebrides (1), Australia (1), and 
Samoa ( 1 ) . 
Intraobtutus, sect. nov. Stigma centripet- 
alum anguste ellipticum in latere proximo prae- 
rupto carpelli truncati affixa; carpellis plurimis 
et adnatis; plantis masculis incognitis. 
Stigma centripetal, narrowly elliptic, borne 
on the proximal steep inner side of the truncate 
carpel apex; carpels 7, adnate 6/7 way; stami- 
nate plant unknown. 
HOLOTYPUS: Pandanus Parkinsonii Martelli, 
Webbia 4(2): 417, t. 38, figs. 14-15, 1914. It 
occurs in the Bismarck Archipelago. 
Lateriobtutus, sect. nov. Carpellis plurimis 
adnatis, stigmatibus carpellorum altrorum ad 
carpellam marginalem dirigitis vel ad lineam 
marginalem carpellorum dirigitis. Ei carpelli ob- 
tuti quam altros breviori. Arbori masculi in- 
cogniti. 
Carpels several, adnate, the stigma of all the 
others facing one marginal or the line of several 
marginal ones at the proximal edge. The focal 
carpels shorter than the others. Staminate trees 
unknown. 
HOLOTYPUS: Pandanus biakensis St. John, 
from Biak, Netherlands New Guinea, H. St. John 
26,142, the description of which is appended at 
the end of this paper. 
Mammillarisia, sect. nov. Phalangibus in 
apice angustatis truncatis lobatisque, apicibus 
carpellis pluribus approximatis depresse conicis 
vel pyramidatis, columnis staminalis diffuse ram- 
osis filamentis brevibus ferrentibus. 
Phalanges narrowed to a truncate apex, lobed; 
apices of the several carpels depressed conic or 
pyramidal, crowded; staminal columns diffusely 
branched, bearing short filaments. 
HOLOTYPUS: P. mammillaris Martelli and 
Pic.-Ser., Inst. Sci. Madagascar, Mem., ser. B, 
3(1): 47-51, fig. 6, a-e, 1951. 
The section contains 26 species, including: 
from west Africa, P. thomensis; from Madagas- 
car, P. bipyramidatus , P. diffusus, P. Karaka, P. 
mammillaris, P. Pervilleanus, P. sambiranensis , 
P . saxatilis, and P. vanDamii; from Reunion, 
P. sylvestris; from Mauritius, P. drupaceus, and 
P. glaucocephalus; from the Philippines, P. acla- 
dus, P. biliranensis, P. camarinensis , P. luzon- 
ensis, P. Martellii, P. nobilis, P. panayensis, P. 
radicans, P. sibuyanensis, P. umbonatus, and P. 
urdatenensis ; from Palau, P. Kanehirae; from 
New Guinea, P. Kaernbachii; and from Aus- 
tralia, P. Dammannii. 
Martillidendron Pic.-Ser., Inst. Sci. Madagas- 
car, Mem., ser. B, 3(1): 20, 1951. Type: P. 
androcephalanthus Martelli. The section contains 
three species from Madagascar. 
Multispina Fagerlind, Svensk Bot. Tidskr. 
34: 112-113, 1940. Type: P. multif meatus 
Fagerlind, from Java, the only species. 
Pandanus. Type: P. odoratissimus L. f., of 
Ceylon. A synonym is the sectional name Keura 
(Forsk.) Kurz, Jour. Bot. Brit, and For. 5: 105, 
1867; Asiat. Soc Bengal, Jour. 38(2) : 147-150, 
1869; genus Keura Forsk., FI. Aegypt.-Arab. 95, 
122, 172, 1775, a familiar and long-used name. 
Since this section contains the type species of 
the genus, it is now mandatory to use the same 
name as the name of this section, in compliance 
with Article 22 of the 1956 International Code 
of Botanical Nomenclature. At first sight Solms 
seems to typify Pandanus in his monograph 
(Linnaea 42: 3, 1878): "Typus P. Kurziani,” 
but he does the same for several other species. 
Those are the ones which he chose to use to 
provide group names for groups of species ( sec- 
tions of other taxonomists ) . These are not valid 
subgeneric or sectional names, being binomials 
for collective species. On p. 77 he says: "P. 
odoratissimus L. fil. Nomen delendum omnes 
fere tunc descriptas Pandani species amplectens.” 
Thus Solms rejected the holotype of the mono- 
typic genus on the basis of confusions and 
misdeterminations by subsequent botanists. His 
rejection is unjustified. 
At the moment, the section contains 169 
species. A few of these occur in Africa and in 
the Indian Ocean: east Africa (6 species), 
Comore Is. (1), Madagascar (1), Mauritius 
( 1 ) , Seychelles (2), Ceylon ( 1 ) , India (2), 
Nicobar Is. (2), Andaman Is. ( 1 ) , but the great 
bulk of the species which remain (151) occur 
in the tropical Pacific, from its western shore 
