26 Sargant. — Theory of the Origin of Monocotyledons 
of transition indicated in Diagram V. The cotyledon pos- 
sesses three bundles throughout, and they are continued 
downwards into a triarch root formed according to Van 
Tieghem’s type i. I have not so far hit on any intermediate 
form which would connect this structure with that of Tulipa 
and Lilium . The anatomy of the seedling, however, may 
possibly be modified by the very early transformation of the 
first leaf into a dropper (Irmisch). This occurs also in both 
species of Tulipa . 
3. Tribe Asphodeleae. 
In this tribe I have examined eleven species belonging to 
eight genera. There is no definite symmetry characteristic 
M 
Diagram VI. 
of the tribe as in Tulipeae, but, with the exception of Bidbine 
annua , the vascular structure of all the species examined can 
be linked with great probability to that of Anemarrhena 
(Diagram VI). 
The seedling of Anemarrhena asphodeloides has been 
described elsewhere in detail (Sargant, 34). It resembles that 
of Alhuca , except in possessing no lateral bundles in the 
cotyledon, and in the regular formation of a tetrarch root. 
Asphodeline liburnica. The cotyledon contains two massive 
bundles throughout, which are quite distinct but orientated in 
the characteristic way — that is, with their protoxylem groups 
directed towards each other. The symmetry of the root- 
stele is sometimes derived from the cotyledonary traces only, 
B C 
A 
