founded on the Structure of their Seedlings . 29 
contains two bundles which are quite distinct from each other 
though close together. Near the base of the sheath the 
bundles so far unite that they have a common protoxylem 
group, and they then form a typical double bundle such as we 
find in Allium (cf. PL V, Figs. 2 and 3). 
The plumular bundles unite to form a single trace which 
then divides into two branches, Pl l9 PI 2 , and these advance to 
meet the double cotyledonary trace M 1 + M 2 at the first 
node (PI. IV, Fig. 2). The protoxylem of the double trace 
has formed three groups already (px h px 2 , px 3 in Fig. 2). 
They represent two branches from the xylem of M x and two 
from that of M 2 . The single group px x is formed by the 
early fusion of two adjacent branches : px, belongs to M 2 and 
px 3 , to M 2 . 
A little lower down, the stele threatens to become diarch 
(PI. IV, Fig. 3). The phloem from M x has met half the 
plumular phloem on one side of the stele, and that from 
M 2 has met the other half on the opposite side. The proto- 
xylem groups within the phloem (px 2) px 3 , in Fig. 3) seem 
about to be obliterated. But they recover themselves later, 
and the root becomes typically tetrarch (PI. IV, Fig. 4). 
Two phloem groups in the tetrarch root are derived from 
the plumule and two from the cotyledon. One protoxylem 
group (px v Fig. 4) is purely cotyledonary and one purely 
plumular [px With regard to the two remaining groups 
of protoxylem, they appear in the young seedlings I have 
examined to be derived from the cotyledon, but it is very 
possible that in older seedlings they would be found to receive 
elements from the plumular traces. If so, the transition 
would be precisely comparable with that of Muscari armenai - 
cum , seedling A 5 (Diagram II). 
Anthericum Liliago. The two bundles of the cotyledon are 
more reduced than those of Chlorogalum. They are united 
by a common protoxylem group throughout their course, and 
in fact appear as a typical double bundle in the upper as well 
as the lower part of the cotyledon. 
