founded on the S true hire of their Seedlings. 43 
For some time I was tempted to look on Doryanthes as 
a genus in which the seedling possessed characters more 
primitive in some respects than those of Anemarrhena. The 
two pairs of bundles, though distinct throughout, might be 
identified with the two massive bundles in the cotyledon of 
Anejnarrhena , each of which behaves like a double bundle 
during the process of transition. The fact that the two pairs 
of cotyledonary traces enter the plumular stele at two distinct 
points suggested at first their origin from distinct members, 
but the arrangement may be merely adaptive to secure greater 
mechanical stability at the insertion of the cotyledon. 
Until the seedlings of allied forms have been examined 
there can be no further comparative evidence concerning the 
origin of the peculiar vascular structure of Doryanthes. The 
seedlings of Agave and Bravoa — two genera nearly related to 
Doryanthes by their mature characters — are, as we have seen, 
designed on a Liliaceous model. I am inclined to believe 
the vascular symmetry of the Doryanthes seedling to be 
derived from that of some form resembling Agave. There 
are two reasons against supposing it primitive. 
In the first place the seedling of Doryanthes is, both 
anatomically and in external form, of the shrubby or arbo- 
rescent type, which is, as a rule, much modified in response 
to its environment. In the second, the floral structure of 
Amaryllids is clearly derived from the Liliaceous type, and 
we are therefore less likely to find a primitive form among 
them than within the Liliaceae. 
Iridaceae. 
Four species representing two genera have been examined 
from this family. 
The three species of Iris, I. sibirica , I. Boissieri , and an 
unnamed species from China, agree in possessing a single 
massive bundle which runs the whole length of the cotyledon, 
and opens out into a double bundle of characteristic form near 
the base of the sheath. Plumular traces take part in the 
transition to a root-structure, and the root is tetrarch ( Iris sp. 
