founded on the Structure of their Seedlings . 13 
of the two main bundles themselves is affected by these 
irregularities, but in some specimens of both species they 
follow the Anemarrhena type exactly, and in almost all 
they clearly begin on that plan, though it is not pursued 
throughout the transition. 
Species in which the transition presents such irregularities 
cannot be fully described here : they are mentioned because 
in individuals of both species a modification of the Anemar- 
rhena transition is found which reappears elsewhere. 
In this variant the phloem of each main bundle divides 
into three groups, of which the median one remains in situ 
A B 
C 
and is continued downwards unchanged. The right-hand 
branch unites with the adjacent branch from the other bundle, 
and a similar fusion occurs on the opposite side of the stele. 
The protoxylem of each main bundle divides into two groups, 
each of which becomes external as it takes up its position on 
one side or other of the median phloem group. The root 
is of course tetrarch (Diagram I). 
The examination of the transitional region in six seedlings 
of Galtonia candicans has convinced me that the type of 
transition just described is derived from type 4 [Anemarrhena). 
It stands in precisely the same relation to that type as Van 
Tieghem’s type 3 to his type i 1 , and for convenience I shall 
refer to it as type 5. 
The genera Hyacinthus and Muscari include a number 
1 Traite de Botanique, ed. ii, 1891, vol. i, p. 782. 
