Dale. — Observations on Gymnoascaceae. 591 
Later they place Eidamella amongst the Gymnoasceae 1 i 
near to Myxotrichum. 
With regard to Endomyces decipiens , no sexual organs 
have been found, but the life-history of G. candidus tends to 
confirm the views of Matruchot and Dassonville. In neither 
G. candidus nor Endomyces decipiens are there any thickened 
hyphae, but in both the asci are completely naked and borne 
on delicate colourless hyphae. In both the mycelium breaks 
up into oidia 2 . The life-history of Endomyces would probably 
repay reinvestigation, with a view to ascertaining the presence 
or absence of sexual organs before the production of asci. The 
chief difference between these two species is that in Endomyces 
decipiens each ascus contains only four ascospores, whereas in 
Gymnoascus candidus there are eight spores in each ascus. 
Van Tieghem 3 also compares Gymnoascus with Hypomyces 
(Endomyces) on the one hand, and on the other with Peni- 
cillium. 
Boudier 4 , as well as Matruchot and Dassonville, regards the 
Gymnoascaceae as having close affinities with Onygena. Indeed, 
he considers that the Gymnoascaceae do not differ essentially 
from the sessile species of Onygena. 
Matruchot and Dassonville claim that Marshall Ward’s 
recent work on Onygena 5 confirms their views as to the relation 
between the Gymnoascaceae and the Onygenaceae. Though 
no definite coil was seen by this author, the resemblances 
between the two families are very strong. In both the ascus 
formation is preceded by a coil, and the asci and ascospores 
develop in the same way. In both families there are chlamydo- 
spores ; in both pyriform swellings and cyst-like swellings at 
the ends of the hyphae occur in the vegetative mycelium. But 
since definite sexual organs are unknown in Onygena , its exact 
systematic position is uncertain. 
1 loc. cit. (1), p. 128. 
2 loc. cit., p. 155, Fig. 135. 
3 loc. cit. (1), p. 161. 
4 Description de deux nouvelles especes de Gymnoascus de France. Bull. Soc. 
Myc., tom. viii, p. 43 (1892). 
5 loc. cit. 
