Notes . 
627 
the micropyle. The existence of these chambers is indicated on the 
outside surface of the seed by the presence of nine little ridges dis- 
posed like the rays of a star around the micropyle, but dying out 
almost at once. These ridges over-lie the partitions of the chambered 
portion of the integument just as do the stigmatic bands the septa of 
a poppy capsule. The whole structure from within is like a fluted 
dome or canopy, the convexities of which correspond to the chambers, 
and actually engage with broad low grooves on the surface of the wall 
of the pollen-chamber. 
The vascular system of the seed enters as a single supply-bundle at 
the chalazal papilla, and branches, a little below the base of the macro- 
spore, into nine radially-running bundles. Each of these bundles 
passes, without further branching, to the apex of the seed, running 
outside the macrospore and a little distance below the surface. At 
the canopy the bundles enter the chambers and end at the tips. 
Lagenostoma Lomaxi was thus a seed or seed-like structure detached 
as a whole and containing pollen-grains in the remarkable cleft-like 
pollen-chamber ; the integument in its free part, when compared with 
that of Williamson's Lagenostoma physoides , suggests a number of 
originally free arms or processes that have become ’laterally fused into 
a complex chambered organ. 
The seed, Z. Lomaxi , is in some cases still attached to its pedicel 1 ; 
the great peculiarity of this seed, as compared with other members of 
the genus, is that when young, and sometimes even at maturity, it is 
found enclosed in an envelope or cupule, springing from the pedicel 
just below the base of the seed, and extending above the micropyle — 
at least in young specimens. The cupule appears to have been 
ribbed below, and deeply lobed in its upper part ; in form it may 
be roughly compared to the husk of a hazel-nut — of course on a very 
small scale. 
The pedicel and cupule bear numerous capitate glands, of which 
some are practically sessile, others shortly stalked, while in others 
again the stalk is of considerable length. The head, or secreting 
portion of the gland, which is spherical in form, is almost invariably 
empty, only the multicellular wall persisting. The tissue of the stalk 
of the gland, consisting of many layers of cells, is preserved, though 
in a somewhat disorganized state. 
These cupular glands present the closest agreement in size, form, 
1 Cf. Williamson, loc. cit., Pt. VIII, Fig. 68 (Z. ovoides). 
U U 
