628 
Notes . 
and structure with the glands which occur on the vegetative organs 
of Lyginodendron Oldhamium 1 , and which are especially abundant on 
the particular form of that plant found in association with Lagenostoma 
Lomaxi. Both on petiole and cupule the majority of the glands are 
short, those which are not sessile being commonly about 0-4 mm. in 
height. Long-stalked glands, exceeding a millimetre in height, some- 
times occur both on the vegetative organs and on the cupule. The 
dimensions of the head of the gland agree exactly on cupule and 
petiole, the diameter averaging about 0-2 mm. in each case. In both, 
the stalk is usually somewhat narrower than the head, except at the 
base, where it is often considerably enlarged. On the stem, as might 
be expected, the glands are usually somewhat larger than on petiole 
or cupule. 
As. a rule, the structure of the glands on the vegetative organs is 
well preserved, the secretory tissue in the head being perfect. But 
occasionally the vegetative glands are found in the same state of 
preservation as those on the cupule, with the head hollow, owing to 
disappearance of the secretory mass. Where we thus have the two 
organs in a corresponding state of preservation, the agreement between 
the vegetative glands of Lyginodendron and those on the cupule of 
Lagenostoma Lomaxi is found to be exact. 
There is no other known plant from the Coal-measures with glands 
at all similar to those described, nor is it likely that any unknown 
Gymnosperm should so exactly resemble Lyginodendron in these 
characters. On the ground, then, of the glandular structure we 
are led to the conclusion that the seed Lagenostoma Lomaxi can 
have belonged to no other plant than Lyginodendron Oldhamium, and 
more particularly to the glandular form of that type with which the 
seed is associated. 
The state of preservation of the glands and of the cupule as a whole, 
indicates clearly that this organ, as we find it, was in an effete condition, 
having, no doubt, already discharged its functions while the seed which 
it protected was still quite young. 
The vascular system of the cupule was well developed, and is very 
fairly preserved. A number of bundles branched off from the main 
strand of the pedicel, and traversed the cupule throughout its whole 
1 It has long been realized that the name Lyginodendron Oldhamium charac- 
terizes a type rather than a species. It is probable that the very glandular form 
occurring at Dulesgate may deserve specific rank. 
