672 Campbell. — Studies on the Araceae . 
an egg-cell. The other four were at the antipodal end of the 
sac, but there were no clearly indicated antipodal cells, nor 
was there, at this stage, any indication of polar nuclei. 
A second embryo-sac, containing two nuclei, accompanied 
this one. 
Fig. 23, which shows an embryo-sac taken from the 
Jamaican specimens, differs from any cases found in the 
material collected at Kew. At the micropylar end of the sac 
was a well-marked egg-apparatus, consisting of two clearly 
defined synergidae and an egg-cell. At the chalazal end 
were two large nuclei surrounded by a mass of granular 
cytoplasm, and evidently in the early prophases of division. 
It would certainly seem that in this case no polar nuclei could 
be developed, and from older specimens from the same plant 
(Fig. 24) it looks very much as if the basal nuclei assumed at 
once the role of endosperm nuclei, proper antipodal cells 
being quite suppressed. 
The specimen figured as Fig. 18 showed ten nuclei. Four of 
these were at the base of the sac, where there was developed 
a group of four cells, one of which projected into the cavity of 
the sac, and may have been the egg-cell, but it is possible 
that this group of four cells may have been antipodal cells. 
Nearly opposite these were six free nuclei, but no further 
differentiation could be discerned, and what the further history 
of the sac would have been could only be conjectured. 
In the embryo-sac shown in Fig. 12 there were twelve 
nuclei. In this case the pollen-tube could be seen, but it was 
not certain which were the generative nuclei. Above the sac 
was a second one, in which were two conspicuous nuclei, 
which may possibly have been the generative nuclei derived 
from the pollen-tube, but they were much more probably the 
nuclei of the second embryo-sac. The apex of the lower 
embryo-sac contained no nuclei. At the lower end were four 
nuclei ( b ) with delicate membranes between them, and prob- 
ably to be considered as antipodal cells. Near them was 
a second group of four cells, which may have represented the 
egg— apparatus, and on the opposite side of the sac were four 
