1J2 Barker . — The Morphology and Development of 
Fig. 2, e). The formation of the septum at this point marks 
off a single cell, composed of the whole of the small hypha 
developed as the result of the growth of the small pro- 
tuberance or new growing point mentioned above. This cell, 
for reasons given later, will be henceforward termed the 
ascogonium , while the original terminal cell of the parent 
hypha, clasped by the ascogonium, will be described as the 
antheridial branch. In those instances in which the asco- 
gonium has been produced at some distance from the apex 
of the parent hypha, that portion, the direction of which 
has been deflected by the growth of the ascogonium, will 
be regarded as the antheridial branch. 
There have now been formed two organs, the ascogonium 
and the antheridial branch, the subsequent behaviour of which 
leads to the production of the ascocarp. They must there- 
fore be regarded as the archicarp. Both of them at this period 
are filled with derise semi-transparent protoplasm, devoid of 
vacuoles but containing a few bright granules. 
The next step in the development of the ascocarp consists 
in a fusion between the two organs. The fusion takes place 
in most cases at the tip of the ascogonium, between that 
portion of it and the part of the antheridial branch in its 
immediate neighbourhood (see Fig. 2,/). Occasionally, how- 
ever, fusion takes place a little distance below the tip of the 
ascogonium (see Fig. 8 ). The fusion appears to take place in 
the following manner. A small papillar outgrowth is developed 
on the antheridial branch at the point, where fusion eventu- 
ally takes place, between that organ and the closely applied 
ascogonium. Solution of the walls then occurs at the point 
of contact between the papilla and the ascogonium, the proto- 
plasm of the two organs becoming continuous. There are 
two reasons for supposing the fusion to take place in this 
manner. Firstly, when the fused portion has become more 
conspicuous, the antheridial branch at the place of fusion is 
slightly swollen on the side in contact with the ascogonium 
and apparently projects into it. That is to say, it is the wall 
of the antheridium that constitutes the wall of the communi- 
