2 1 8 Barker . — The Morphology and Development of 
thickened wall and passing through a long period of rest, as 
is the case with them, it proceeds at once to swell up con- 
siderably and produce ascogenous hyphae. This process 
must be regarded as the germination of the fertilized ‘cell’ 
and the beginning of the sporophyte generation : and there- 
fore corresponds with the germination of the oospore and the 
production of the promycelium of such a form as Phytophthora 
omnivora De Bary. The generation of the sporophyte is 
terminated in the one case by the formation of ascospores in 
asci, and in the other case by the formation of zoospores 
in sporangia. The germination of these spores in the respec- 
tive instances gives rise to the gametophyte. A difference is 
shown in the time of germination ; the zoospores, which are 
naked masses of protoplasm, germinating immediately after 
coming to rest and clothing themselves with a cell- wall, while 
the ascospores, which are thick-walled resistant bodies, are 
specially prepared to pass through a period of rest during 
unfavourable external conditions before germination at a 
suitable time. Thus the functions of the oospores are not 
assumed by the fertilized 4 central cell, 5 but are passed on to 
the ascospores. The shifting of the period of the resting 
stage of the organism here indicated suffices to explain the 
differences of organization of the female ‘ cells ’ in the respec- 
tive cases, and renders the parallel behaviour of the respective 
Fungi as regards the course of their life histories much closer 
than appears structurally. Leaving out of the question the 
differences which may be taken as arising naturally from the 
delegation of the function of hibernating to different structures 
in the two cases, there is a remarkable degree of resemblance 
in the manner and the course of the reproductive processes of 
these Fungi, so widely different in habit and in the structure 
of the mycelium. It is certainly sufficiently marked to make 
the idea of relationship likely. 
A further point of resemblance between Monascus and the 
Oomycetes is the nuclear behaviour during the reproductive 
processes. It has been shown that in Monascus several nuclei 
from the antheridium probably pass into the ascogonium and 
