On the Development of the Sexual Organs 
and Fertilization in Picea excelsa. 
BY 
K. MIYAKE. 
With Plates XVI and XVII. 
Introductory. 
I N spite of the activity which has of recent years been 
manifested in the study of the fertilization and embryogeny 
of the Abietineae, the genus Picea has been seemingly neglected. 
In fact, our knowledge on the cytology of the sexual repro- 
duction of Picea remained almost stationary for nearly ten 
years. In the light of the recent studies on the other members 
of the Abietineae, it is highly desirable that similar investiga- 
tions should be made in Picea. It was with this object that 
the work described in the present paper was undertaken. 
But few investigators have hitherto studied the development 
of the gametophytes and fertilization in Picea. Hofmeister 
(’51, ’58, ’62), who made the first careful and extensive studies 
on the life-history of the Conifers, has only incidentally 
referred to Picea. 
Strasburger (’69) in his first work on the fertilization of the 
Conifers described the structure of the archegonium and the 
formation of the proembryo in Picea excelsa (P. vulgaris ) with 
several figures. He traced the pollen-tube into the arche- 
[Annals of Botany, Vol. XVII. No. LXVI. March, 1903.] 
