734 G wynne- Vaughan. — Observations on the 
to go together, and they both come under Bower’s division 
of Gradatae. It seems probable also that the genus Loxsoma 
should be regarded as more nearly allied to these two genera 
than to any others. The development of the sorus in Lepto - 
lepia and Saccoloma is not known, but Hypolepis belongs to 
the Mixtae. According to the anatomy Leptolepia goes with 
the Microlepias, but it may prove that a distinction should be 
made between Saccoloma and Hypolepis and the rest of the 
group, and also between each other. As a secondary charac- 
ter of the Dennstaedtiinae Prantl states that, except in Sacco- 
loma , hairs are present instead of paleae throughout the group. 
So far as the species that I examined are concerned this 
certainly holds true. 
The Lindsaya- type of primitive stele has been found in all 
the species of that order hitherto examined, except in L. retusa 
and L. cultrata , which possess solenosteles. The same type 
of stele is also characteristic of the sections Odontoloma and 
Stenoloma , which are placed by Hooker in the genus Davallia. 
The anatomy, therefore, agrees with H. Christ in removing 
these two groups from Davallia and placing them among the 
Lindsayas. J. Smith also places Odontoloma in his tribe 
Lindsaeeae. The section Stenolojna he removes from the 
Davallias, but places it in his tribe Saccolomeae under the 
synonym of Odontosoria. It should be mentioned that 
Davallia ( Stenoloma :) aculeata is somewhat exceptional, in that 
it exhibits a type of solenostely peculiar to itself. H. Christ 
has given it a section of its own : Lindsayopsis. It has 
already been stated that according to the anatomy Davallia 
{Microlepia) pimiata should be included with the Lindsayas 
rather than with the Microlepias. The most primitive genera 
in Prantl’s second tribe, the Asplenieae, were considered by 
him to be Lindsaya , Lindsayopsis , Wibelia {Davallia pinnata), 
Odontosoria {Stenoloma) and Davallia. If the last genus, 
Davallia , be excluded he finds here again support from the 
anatomy, since all the species in the four remaining genera 
possess a primitive vascular system, and, with three exceptions 
only, they all possess the same type of stele. Finally they all 
