New Pogonophora — -H artman 
543 
mm. The body is nearly uniform in width but 
is widest in the mesosomal region where it meas- 
ures 0.065 mm.; the postannular region grad- 
ually becomes slenderer and tapers posteriorly. 
The tentacle is less than half as wide as the body 
and extends forward for a length nearly two and 
a half times that of the mesosomal length. Its 
insertion marks the ventral side of the body as 
herein interpreted, as well as the posterior end 
of the first body region or protosoma. The three 
body regions are not sharply separated from one 
another. The separation between the first and 
second regions is behind the tentacular insertion 
and is vaguely indicated by a faint line extend- 
ing obliquely forward around the body to the 
middorsum. The second region, or mesosoma, 
is about four times longer and is separated from 
the third region, or metasoma, by a transverse 
groove located just in front of the paired gland- 
ular papillae (see Figs. 5, 6). The third region 
is much the longest and comprises most of the 
length of the body. 
The tentacle is extended forward nearly 
straight in some specimens and is loosely to 
closely coiled in others, or only a short distal end 
is coiled. Most of the length has lateral, filiform 
branches or pinnules, inserted in two nearly reg- 
ular rows along the ventrolateral edge (Fig. 4); 
those nearer the distal end are more dispersed, 
whereas those nearer the base (Fig. 3 ) are closer 
together and tend to be curled toward the mid- 
ventral line. Each pinnule is very slender and 
longer than the tentacle is thick. These features 
agree with those described for other species of 
the genus having pinnules; only two ( S . inermis 
Ivanov and S. atlanticum Southward and South- 
ward) are said to lack them. 
The protosoma or anteriormost region of the 
body is short, subconical, tapers forward, and 
lacks markings; its posterior end is indicated on 
the ventral side of the body by the insertion of 
the tentacle. This fusion is firm, so that the two 
are not easily severed. A mouth, if present, would 
be expected at the posterior base of this ten- 
tacle, for which the principal function seems to 
be that of food gathering. I have seen no oral 
aperture or any indication of an alimentary tube 
but would not exclude the possibility of a length- 
wise transport tube of some kind in an animal 
of such great length. 
The second body region, the mesosoma, is 
considerably longer than the first and is cylindri- 
cal in shape; near its anterior end it is obliquely 
crossed by the bridle or frenulum (Ivanov) or 
girdle (some other authors). It is separated 
from the third region, or metasoma, by a trans- 
verse groove. A pair of circular pores is located 
at the sides, just behind the frenulum (Fig. 6); 
they are believed to be the external apertures of 
the coelomoducts for the second segment. 
The anterior end of the metasoma is charac- 
terized by the presence of ventrolateral paired 
glandular papillae; they are so arranged that the 
two members of a pair are separated from suc- 
cessive ones by a space about equal to that of 
the distance between the brown bars on the 
tube. It thus seems obvious that these papillar 
glands are directly concerned with tube secre- 
tion. This region is followed by the much longer 
goniadial region, characterized by sparsely and 
irregularly dispersed epithelial papillae. About 
halfway along the length of the body two trans- 
verse series of uncinial rows indicate the posi- 
tion of the annular region; for this reason the 
goniadial region in front is called the preannu- 
lar, and the same one behind is the postannular, 
region. 
The annulae are of particular interest because 
they bear hard, chitinized platelets, or unicini, 
in transverse series. Those of the anterior row 
are incomplete and number about 38, while 
those in the posterior row are more numerous, 
numbering about 44. All uncini are of one kind, 
and terminate distally in a single row of denti- 
cles (Fig. 8), numbering 9—13- Each uncinus is 
distally recurved so that those of successive un- 
cini tend to overlap. Seen from the end the den- 
ticles form an arcuate row (Fig. 9) . The basal or 
embedded stems are not chitinized but taper 
rapidly to very tenuous strands, and are so pro- 
longed that all those of one transverse series are 
brought together as a fascicle and carried ob- 
liquely forward to attach to the body wall. They 
may function to anchor the animal within the 
tube at selected places. The comparable uncini 
in some other species, S. atlanticum and S. iner- 
mis, have been shown as having multiple rows 
of crenulations (see Southward and Southward, 
1958: 629, 631). 
S. veleronis was recovered from a quantitative 
sample covering a surface area less than a half 
