Page 64: Revision of Pandanus, 7. Papua and the Solomons — S t. John 
579 
1- 3 in a spike and 11.5-14 cm. long, 13.5-14 
cm. in diameter; drupes 35-40 mm. long, 6-7 
mm. wide; pileus tapering into the style; and 
the endocarp in the lower Va. 
The new species is named in honor of its col- 
lector, Mr. J. A. R. Anderson, Forest Research 
Officer of the Sarawak government. 
Pandanus fruticosus sp. nov. (sect. Acro- 
stigma) 
Figs. 35-36 
DIAGNOSIS HOLOTYPI: Frutex 15.5 m. altus 
ramosus, ramulis foliosis 6-9 cm. diametro, cor- 
tice rubrescenti-brunneo lucido cum cicatricibus 
pallidis cinctis, foliis 51-58 cm. longis 8-10 
mm. latis subcoriaceis ligulatis M-formatis longe 
diminuentibus in apice subulato 10 cm. ex apice 
3.5 mm. lato, marginibus in basi integris sed ex 
4 cm. marginibus cum dentibus validis subu- 
latis 0.5-1. 3 mm. longis 1-3 mm. distantibus 
nigris in 45° adscendentibus, midnervio exar- 
mato, marginibus in sectio mediale cum serris 
nigris 0.5 mm. longis 1-2 mm. distantibus, mid- 
nervio infra cum serris nigris 2-6 mm. longis 
2- 6 mm. distantibus, circa apicem marginibus 
et midnervio infra cum serris acutis 0.4-0. 5 
mm. longis 0.5-1. 5 mm. distantibus, inflores- 
centia foeminea solitaria terminale cum bracteis 
plurimis 16 mm. latis amplectentibus, syncarpio 
subsessile 3 cm. longo 2 cm. diametro ellip- 
soideo, drupis multis 9-10 mm. longis 2-2.5 
mm. diametro fusiformis, corpore 4-5 mm. 
longo, pileo obliquiter conico 4 (-5) mm. 
longo ad stylum tenuato, stylo subulato cum 
linea stigmata proxima 2.5-3 mm. longa, endo- 
carp io supramediale lateribus 0.1 mm. crassis 
pallidis, semine 3.5 mm. longo 1.6 mm. diam- 
etro late elliptico, mesocarpio infero cavernoso. 
DESCRIPTION OF HOLOTYPE: Shrub 15.5 m. 
tall, forking; leafy branchlets 6-9 cm. in diam- 
eter, the bark reddish brown, shining, ringed by 
pale leaf scars; leaves 51-58 cm. long, 8-10 
mm. wide, subcoriaceous, ligulate, M-shaped in 
cross section, long tapering to a subulate apex 
which at 10 cm. back from the tip is 3.5 mm. 
wide, margins at the very base entire, but from 
about 4 cm. up the margins with stout subu- 
late teeth, 0.5-1. 3 mm. long, 1-3 mm. apart, 
wholly black, ascending at 45°, the midrib un- 
armed; at the midsection the margins black 
serrate, the teeth 0.5 mm. long, 1-2 mm. apart, 
the midrib below appressed, subulate, black ser- 
rate, the teeth 2-6 mm. apart; near the apex the 
margins and midrib below sharply serrate, the 
teeth 0.4-0. 5 mm. long, 0.5-1. 5 mm. apart; 
pistillate inflorescence solitary, terminal, sur- 
rounded by several bracteal leaves 16 mm. wide 
around the spike, but otherwise like the normal 
foliage leaves; syncarp subsessile 3 cm. long, 
2 cm. in diameter, ellipsoid, with numerous 
drupes; drupes 9-10 mm. long, 2-2.5 mm. in 
diameter, fusiform, the body 4-5 mm. long, the 
pileus oblique conic, 4 (-5) mm. long, taper- 
ing into the subulate style which is linear stig- 
matic for 2.5-3 mm. on the proximal side; en- 
docarp from middle to apex of body, the wall 
0.1 mm. thick, pale; seed 3.5 mm. long, 1.6 mm. 
in diameter, broady ellipsoid; lower mesocarp 
cavernous. 
HOLOTYPUS: North Borneo, Keningau Dis- 
trict, 76V2 mi. on path from Tenom to Ranau 
(81/2 mi. NE. of Tambunan), 2,100 ft. alt., 
Aug. 27, 1954, G. H. S. Wood & J. Wyatt-Smith 
A 4,4.30 ( bri ) . 
DISCUSSION: The new species P. fruticosus is 
in the section Aero stigma. Its nearest relative is 
the Malayan species P. aurantiacus Ridl. which 
has the leaves 3.8 cm. wide; inflorescence of 5 
heads; syncarp 6.2 cm. long, 2.5 cm. in diam- 
eter; drupes 18 mm. long; and the endocarp 
12 mm. long. On the other hand, P. fruticosus 
has leaves 8-10 mm. wide; the syncarp solitary; 
drupes 9-10 mm. long; and the endocarp 3.7 
mm. long. 
The new epithet is the Latin adjective, fruti- 
cosus, shrubby, given with reference to the plant 
habit. 
Pandanus ruber sp. nov. (sect. Micro stigma) 
Figs. 37-38 
DIAGNOSIS HOLOTYPI: Arbor 3-5 m. alta 
ramosa, trunco cum aculeis adscendentibus ar- 
mato, radicibus fulturosis paucis aculeatis, foliis 
172 cm. longis 10.2 cm. latis coriaceis in sectio 
M-formatis ligulatis acutis in sectio 10 cm. ex 
apice 4 cm. latis in base integris sed ex puncto 
9 cm. e base cum aculeis 1-1.6 mm. longis 2-7 
mm. distantibus salientibus stramineis, mid- 
nervio inerme, in sectio mediale marginibus 
cum aculeis 1-1.5 mm. longis 3-6 mm. distanti- 
