Page 68: Revision of Pandanus, 7. Papua and the Solomons — S t. John 
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bus proxime adpresse-adscendentibus, midnervio 
cum aculeis 0.3-1 mm. longis 3-11 mm. distanti- 
bus valide aciculatis adpresse adscendentibus 
subbrunneis, proxime apice marginibus unice- 
vel duplo-serratis serris 1.5-2. 1 mm. longis 2-5 
mm. distantibus stramineis, supra nervis secun- 
dariis cum aculeis paucis eis 0.8-1 mm. longis, 
infra nervo mediale saliente et cum aculeis si- 
multantibus 1.5-2 mm. longis 3-10 mm. distan- 
tibus, syncarpiis solitariis plerumque 42 cm. 
longis 10.5 cm. diametro cylindricis sed sub- 
triangulatis, receptaculo aurantiaco-rubro, dr ti- 
pis numerosissimis in sectio mediale 13-15 mm. 
longis 3-4.5 mm. latis 2.5-3 mm. crassis unicar- 
pellatis obscure rubris 4-6-angulatis parte Va 
supera libera, pileo 3-4 mm. longo anguste 
pyramidale asymmetrico et arcuato corpore ob- 
longo in base subcuneato, stigmatibus 0.8-1. 5 
mm. longis et latis suborbicularibus brunneis 
verticalibus proximalibus infra petaso terminale, 
endocarpio basale 9 mm. longo osseoso obscure 
brunneo lucido 0.4-0. 5 mm. crasso, mesocarpio 
apicale caverno unico. 
DIAGNOSIS OF holotype: Tree, 3-5 m. tall, 
branched; trunk and branches armed with up- 
turned prickles; prop roots few, prickly; leaves 
172 cm. long, 10.2 cm, wide, coriaceous, M- 
shaped in cross section, ligulate, acute, 4 cm. 
wide at 10 cm. from the tip, at the very base the 
margins entire, but from 9 cm. up prickly, the 
prickles 1-1.6 mm. long, 2-7 mm. apart, spread- 
ing, straw-colored, the nearby midrib unarmed; 
at the middle point the margins with prickles 
1-1.5 mm. long, 3-6 mm. apart, closely ap- 
pressed ascending, those of the nearby midrib 
0.3-1 mm. long, 3-11 mm. apart, stout acicular 
appressed ascending, brownish near the tip the 
margin simply or doubly subulate-serrate, the 
teeth 1.5-2. 1 mm. long, 2-5 mm. apart, straw- 
colored, above the ribs of the two secondary 
pleats with a few ascending prickles 0.8-1 mm. 
long, below the salient midrib with similar 
prickles 1.5-2 mm. long, 3-10 mm. apart; syn- 
carps solitary, averaging 42 cm. long, 10.5 cm. 
in diameter, cylindric but somewhat 3 -sided; re- 
ceptacle, orange-red; with myriad drupes, those 
at midsection 13-15 mm. long, 3-4.5 mm. wide, 
2.5-3 mm. thick, 1 -celled, dark red, 4-6-angled, 
upper Va free, the pileus 3-4 mm. long, nar- 
rowly pyramidal, asymmetric and arcuate to the 
apex, body almost oblong, but slightly cuneate 
below; stigmas 0.8-1. 5 mm. long and wide, 
suborbicular, brown, vertical on proximal side 
of apex, below the apical cap; endocarp basal, 
9 mm. long, bony, dark brown, shiny, the walls 
0.4-0.5 mm. thick; apical mesocarp a single 
cavern. 
HOLOTYPUS: Papua, Central Division, Bella 
Vista, 1,450 m. alt., planted by the natives in 
gullies and ravines close to their gardens, Nov. 
1933, L. J. Brass 5,463 (bri); and isotype (A). 
DISCUSSION: The closest relative is P. Holl- 
rungii Warb. of Deutsch Neu-Guinea, now Aus- 
tralian New Guinea. It differs by having: the 
leaves 2.5 m. long, 5 cm. wide; syncarp 35 X 
4-7 cm., sessile, spathe-surrounded; drupes 11- 
13 mm. long, 2.5—3 mm. wide, the apex semi- 
orbicular-pyramidal; stigmas apical, subhorizon- 
tal, 1.5 mm. wide; and the endocarp slightly 
submedian. P. ruher differs by having: the leaves 
1.7 m. long, 10.2 cm. wide; syncarp 42 X 10.5 
cm.; drupes 13-15 mm. long, 3-4 mm. wide, 
3 mm. thick, the apex narrowly pyramidal, asym- 
metric and arcuate; stigmas 0.8-1. 5 mm. wide, 
vertical on proximal side below overhanging 
apical cap; and the endocarp basal. 
The collector recorded that, "the leaves of 
mature trees always much damaged by in.- 
sects. . . .” Also that the trees were cultivated. It 
is doubtless also wild in the forest. The several 
similar species with cylindric, red syncarps yield 
on boiling a fruit sauce important in the diet of 
the New Guinea natives. 
This plant was identified as P. conoideus Lam. 
by Merrill and Perry (Journ. Arn. Arb. 20: 175, 
1939). Lamarcks species is similar in its sub- 
terminal stigmas, but the drupes are 16 mm. 
long, the pileus is broader, and the endocarp is 
only submedia, and the leaves are narrower and 
with horrid spines. It is based on P. ceramicus 
Rumph ( 1743) from Amboina, Ceram, and the 
Indonesian islands. 
The epithet, ruber , is the Latin adjective 
meaning red, here chosen because of the red 
color of the fruits. 
Pandanus yuleensis sp. nov. ( sect. Pandanus ) 
Figs. 39-40 
DIAGNOSIS HOLOTYPI: Frutex ad 6.6 m. alta, 
cortice radicibus crassis adscendentibus ad 12 
