Page 32: Revision of Pandanus , 5. Maldives and Seychelles — St. John 
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'6-7-angulosis, suturis lateralibus nullis, sinibus 
apicalibus centralibus 3-4 mm. profundis V- 
formatis (phalangiis geminatis 5 — 5«5 cm. lads 
et carpellis 18-22), carpellis 9-13, apicibus hu- 
militer pyramidalibus eis centralibus valde ob- 
latis Mis marginalibus late distal! truncatis et 
etiam concavis, stigmatibus 2.5-4 mm. longis 
cordatis vel irregulariter ellipticis oblique in- 
clinatis centripetalibus, sutura proximal! V 5 -V 2 
distancia ad fundum extenia, endocarpio paene 
submediali osseoso obscure brunneo marginibus 
lateralibus 2 mm. crassis, seminibus 8-10' mm. 
longis 3=5-5 mm. diametro oblique ellipsoideis, 
rnesocarpio apical! in quaque carpella cavernoso 
cum fibris longitudinalibus et membranis rne- 
dullosis, rnesocarpio basali fibroso. 
DIAGNOSIS OF HOLOTYPE: Tree 6-7 m. tall; 
crown broad ovoid; prop roots present; leaves 
150 cm. long, 6.3 cm. wide, firm subcoriaceous, 
M -shaped in cross-section, ligulate, the apex 
gradually long acuminate, but the tip not seen, 
the margins near the base with prickles 3-4 
mm. long, 12-22 mm. apart, stout subulate, 
broad based, ascending, dark brown tipped; the 
nearby midrib below with prickles 2.5-3 mm. 
long, 11-30 mm. apart, reflexed, stout subulate 
from an enlarged base; at the midsection the 
margins with prickles 2-3 mm. long, 8-22 mm. 
apart, subulate, ascending at 30°; the midrib 
unarmed; near the apex the margins and mid- 
rib below with ascending prickles 0.4-0 .6 mm. 
long, stout subulate, ascending at 45°, those of 
the margin 3-5 mm. apart, but those of the mid- 
rib 5—10 mm. apart; phalanges 4.5-4.9 cm. 
long, 3. 4-4.1 cm. wide, 3-3.7 cm. thick, obovoid 
with the base cuneate and the apex convex, up- 
per 14 free, the sides smooth, shining, brown 
when dry, almost plane, sharply 6-7-angled, 
lateral sutures none; central apical sinuses 3-4 
mm. deep, V-shaped; (phalange twins 5-5.5 
cm. wide and with 18-22 carpels); carpels 9- 
13; carpel apices low pyramidal, the central 
ones strongly oblate, the marginal ones broad 
truncate and even concave distally from the 
stigma; stigmas 2.5-4 mm. long, cordate or ir- 
regularly elliptic, oblique, centripetal; proximal 
suture running l A~Vi way to valley bottom; 
endocarp slightly submedian, bony, dark brown, 
the lateral margins 2 mm. thick; seeds 8-10 
mm. long, 3.5-5 mm. in diameter, obliquely 
ellipsoid; apical mesocarp in each carpel a cavity 
with longitudinal fibers and pithy partitions; 
basal mesocarp fibrous. 
HOLOTYPUS: Maldive Islands: Addu Atoll, 
Hitadu Islet, near shore, common, Oct. 13, 1957, 
W . D. Hartman 4 (Arn. Arb. ) . 
DISCUSSION : P. adduensis is a member of the 
section Pandanus . In that section its close rela- 
tive is an undescribed species from Papua, St. 
John 26,234 , which has the phalange upper 
third free, the apex low convex; carpel apices 
rounded pyramidal, the outer ones with an 
oblique outer plane surface reaching only to the 
stigma; stigmas 1-2 mm. long, ellipsoid; bark 
with papillae developing ascending rootlets. P. 
adduensis has the phalange with the upper 
quarter free, its apex convex; carpel apices low 
pyramidal, the marginal ones broad truncate 
and even concave distally from the stigma; 
stigmas 2.5-4 mm. long, cordate or irregularly 
elliptic; and the bark lacking ascending rootlets. 
The new epithet is made of Addu, the type 
locality; and - ensis , the Latin adjectival place 
ending. 
Pandanus Fosbergii sp. nov. (sect. Pandanus) 
Figs. 13-14 
NOM. VERN.: "karikio.” 
DIAGNOSIS HOLOTYPI: Arbor 6-8 m. aka, 
foliis 3-4 ("ad saltern 4”) m. longis 11-13 cm. 
latis firmis spongioso-coriaceis minime AA-for- 
matis in sectio ligulatis sed ad apicem con- 
strictis in apice (acuto? ), basi amplexicaule 
exarmatis per 10 cm. deinde marginibus grosse 
subulato-serratis serris 3.5-5 mm. longis 10-25 
mm. distantibus in apice brunneis, midnervio 
infra cum serris simulantibus 3 mm. longis 15- 
30 mm. distantibus reflexis, in regione mediali 
marginibus cum serris 3-4 mm. longis 7-17 
mm. distantibus grosse subulatis adpresse ad- 
scendentibus, midnervio infra salient! et cum 
dentis sumulantibus adscendentibus 4-8 cm. dis- 
tantibus, proxima apice marginibus subulato- 
FlG. 13. Pandanus Fosbergii, from holotype. a, Phalange, lateral view, i| 1; b, phalange, longitudinal 
section, X 1*, G phalange, apical view, X 1 \ d, e, stigmas, oblique view, X 4. 
