Page 42: Revision of Pandanus, 5. Maldives and Seychelles — St. John 
341 
anything, April 12, 1956, F. R. Fosberg 36,916 
(US). 
DISCUSSION: P. maldivecus is a member of 
the section Pandanus. Its closest relative seems 
to be P. tectorius Sol. var. sarawakensis Martelli 
of Borneo. This has the phalanges 6-6.5 cm. 
long, the upper Vl free, the sides flat or more 
or less concave; central apical sinuses apparently 
4-5 mm. deep; and the stigmas 2-2.8 mm. long, 
indexed even on the marginal carpels. In con- 
trast, P. maldivecus has the phalanges 4.8-5 .2 
cm. long, upper free, the sides plane and 
curved or shallowly furrowed; central apical 
sinuses 2-3 mm. deep; and the stigmas of the 
marginal carpels distinctly diverging. 
The new specific epithet is taken from the 
geographic name Maidive, to which is added a 
Latin adjectival suffix. 
Pandanus OF THE SEYCHELLES ISLANDS 
Pandanus Balfourii Martelli, Webbia 1: 361— 
362, 1905; 4(1): 7, t. 4, figs. 4-7, 1913 
( sect. Pandanus ) 
Fig. 19 
This species was briefly described, then later 
illustrated by Martelli, from collections from 
the Seychelles made by Horne and by Dupont. 
It is a well-marked species, and its status is not 
in question. However, the collection, data, and 
photos by Dr. W. D. Hartman, made in 1957, 
make possible an expanded and more complete 
description, including stems, leaves, etc. The 
supplementary descriptive details are as follows: 
Tree 5-6 m. tall, and at a point just above 
the roots 15-17.5 cm. in diameter, but below 
the first branches 20-23 cm. in diameter, ap- 
pearing smooth; crown semi orbicular; prop 
roots borne up to 6 dm. from the ground; 
leaves 125-145 cm. long, 4.6-4.S cm. wide, 
firm, strong, subcoriaceous, ligulate, AA-shaped 
in cross-section, the tip long, slender acuminate 
to a subulate tip, this at 10 cm. down only 2 
mm. wide, at 20 cm. only 8 mm. wide, near 
the base the margins with prickles 3-3.5 mm. 
long, 3-10 mm. apart, stout, broad-based subu- 
late, salient ascending, the nearby midrib below 
beginning 12 cm. up with prickles 2.5-3 mm. 
long, 10-25 mm. apart, stout subulate, reflexed; 
at the midsection the margins with prickles 2— 
2.5 mm. long, 2-5 mm. apart, appressed ascend- 
ing and more slenderly subulate, tipped with 
dark brown; the midrib below with prickles 
l. 5-2 mm. long, 7-15 mm. apart, slender subu- 
late, ascending-subappressed; near the apex the 
margins and midrib below with prickles 0.3- 
0.5 mm. long, 2-3 mm. apart, stout subulate, 
broad based, appressed-ascending, brown tip- 
ped; phalanges 102; these 3.7-4 cm. long, 2- 
4.3 cm. wide, 1.6-2. 5 cm. thick, upper 1/5 -Va 
free, apex low convex, sides gently convex or 
plane, shining, pale brown when dry; lateral 
sinuses none; lateral furrows none or slight, 4— 
6- angled, lower half fibrous and fleshy and 
orange; central apical sinuses 1-2 mm. deep, 
shallow, broad V-shaped; carpels 5-10, their 
apices low, rounded pyramidal, green when ripe; 
stigmas 1-2 mm. long, suborbicular to cordate; 
endocarp dark brown, slightly supramedian, the 
lateral margins 1-2 mm. thick. 
SEYCHELLES ISLANDS: Mahe Island, opposite 
Souris Island, group on granite cliffs near shore, 
Nov. 9, 1957, W . D. Hartman 6 (Arn. Arb. ). 
Pandanus Hornei Balf. f., in Baker, FI. Mauritius 
and Seychelles 397, 1877; Martelli, Webbia 
4(1): 17, t. 16, figs. 1-3, 1913 (sect. 
Dauphinensia ) , new sectional placement 
Figs. 20-21 
NOM. vern.: "vacoa parasol.” 
This very distinct species has been known for 
nearly a century. The phalanges were later illus- 
trated by Martelli, but his figures did not clar- 
ify the remarkable stigmatic apparatus. Now, 
the recent collection and data by Dr. W. D. 
Hartman allow the furnishing of a supplemen- 
tary description and more adequate illustrations, 
as follows: 
Trees 15-20 m. tall, and the trunk at 3 m. 
from the ground 15-17.5 cm. in diameter, at 
7- 10 m. up 20-22.5 cm. in diameter, tall 
straight, twice trichotomous, then dichotomous; 
prop roots 2.4-3 m. long, nearly vertical and 
seeming to hug the trunk; leaf (1 seen) 2.25 
m. long, 10 cm. wide, subcoriaceous, gradually 
tapering to the elongate apex (but the tip lost), 
at base 1 -furrowed, further out M -plicate, at 
base the margin entire for 11 cm., then in- 
