40 
selected as dominants on a standing crop basis 
retain this rank when secondary productivities 
can be calculated. Also, we need to be able to 
recognize successional stages in the sublittoral, 
for until we do it is impossible to assume that 
a small contributor to the biomass at one time 
interval is not an essential part of the community 
over a longer time-span. 
PACIFIC SCIENCE, Vol. XXII, January 1968 
REFERENCES 
Pequegnat, W. E. 1964. The epifauna of a 
California siltstone reef. Ecology 45(2) : 272- 
283. 
Turpaeva, E. P. 1957. Food interrelationships 
of dominant species in marine benthic bio- 
coenoses. Trans. Inst. Oceanol. 20:137-148. 
USSR Acad. Sci. Press, Moscow. 
