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PACIFIC SCIENCE, Vol. XXII, January 1968 
ligulate, at the tip narrowing into a short tri- 
gonous subulate apex which 10 cm down is 
6-10 mm wide; near the base beginning at 
16-17 cm the margins with spines 3-5 mm 
long, 4-10 mm apart, subulate from a stout 
base, the lower ones divergent, then the next 
slightly ascending; the midrib below beginning 
at 19 cm with spines 3 mm long, 4-8 mm 
apart, the base broad conic, the apex subulate, 
recurved ; at midsection the margins with prick- 
les 2-3 mm long, 3-10 mm apart, subulate, 
ascending; the midrib below with prickles 1.5- 
2.5 mm long, 2-7 mm apart, subulate, appressed 
ascending; on the subulate apex the margins 
and midrib below with subulate serrulations 
0.2-0. 3 mm long, 0.5-3 mm apart, white; 
lateral branchlets 2-2.5 cm in diameter, brown, 
sharp conic spiny right up to the leaves ; leaves 
1-1.2 m long, 18-22 mm wide, thin, coria- 
ceous, above dark olive-green, below pale green 
and glaucous, gently curved, not breaking across, 
above with a broad V-shaped furrow occupying 
the middle half, the lateral quarters pleated but 
horizontal, near the apex the two pleats sharply 
serrulate above, at midsection with 21 parallel 
secondary veins in each half, no tertiary cross 
veins, blade ligulate, narrowing near the tip to 
a trigonous subulate apex, this 10 cm down 
6- 10 mm wide, the base unarmed and amplex- 
icaul, white, the margins near the base begin- 
ning at 15 cm with prickles 0.5-1. 2 mm long, 
4-10 mm apart, from serrulate to subulate, 
ascending, white, brown-tipped; midrib below 
unarmed to the middle or beyond; at midsec- 
tion the margins with serrulations 0. 2-0.9 mm 
long, 2-4 mm apart, white; the midrib below 
unarmed or with serrulations 0. 1-0.2 mm long, 
7- 12 mm apart, white; on the subulate apex 
the margins and midrib below with serrulations 
0.3-0. 5 mm long, 2-3 mm apart, white; pistil- 
late inflorescence not seen, but reported to be 
"on the small, lateral branchlets, spicate, yel- 
low.” Phalanges 20-22 mm long, 10-15 mm 
wide, 8-11 mm thick, obovoid, compressed, 
4-7 -celled, upper y 3 of body free; styles 6-7 
mm long, acicular, outward arcuate, borne from 
a short conic base ; stigmas 6-7 mm long, linear, 
proximal, brown, papillose; endocarp in upper 
y 3 of body, bony, brown, the lateral walls 1- 
1.5 mm thick, the inner surfaces dark brown, 
smooth, shining; seeds 3.5-5 mm long, ellip- 
soid; apical mesocarp 3 mm long, sparse, with 
strong longitudinal fibers; basal mesocarp fi- 
brous and fleshy. 
holotypus: Malagasy Republika, Andasibe, 
ligne Cap Est, 20 km S of Antalaha, in rem- 
nant of coastal forest, infrequent, 5 m alt, 
6 Sept. 1961, H. St. John 26,571 (bish). 
discussion : P. columnaris is a member of 
the section Acanthostyla, as is its closest rela- 
tive, P. mangokensis Martelli, a species with the 
drupes 12-15 mm long, 7-9 mm wide, the body 
ovate-obpyramidal and 9-11 mm long; styles 
1-3, and 4-5 mm long; endocarp "ad basim 
drupae sita” ; terminal leaves 1.4-1. 6 m long, 
8 cm wide; leaves of lateral branchlets 80 cm 
long, and 1-1.4 cm wide. P . columnaris has 
the drupes 20-22 mm long, 10-15 mm wide, 
the body 14-15 mm long, broad obovoid; 
styles 6-7 mm long; endocarp in the upper y 3 ; 
terminal leaves 2 or more meters long, 24 cm 
wide; leaves of the lateral branchlets 1-1.2 m 
long, and 1.8-2. 2 cm. wide. 
The new epithet is the Latin adjective colum- 
naris, columnar, and is descriptive of the shape 
of the crown of the tree. 
Pandanus pluriloculatus sp. nov. (sect. Acan- 
thostyla) 
Figs. 248, 249, 250^, b. 
nom. VERN.: "fandran.” 
diagnosis holotypi: Arbor 22 m alta 30 cm dia- 
metro, cortice obscure brunneo cum spinis conicis, 
radicibus fulturosis plerumque nullis, corona anguste 
cylindrica in apice cum spiris tribus foliarum gran- 
dium sed infra cum ramulis multis brevibus aequa- 
libus cum foliis multis angustioribus et fructubus, 
foliis terminalibus 1.35-1.6 m longis 10-14 cm latis 
crassis coriaceis V-sulcatis et proxima apicem 2-plicatis 
in sectione mediali cum 79 nervis parallelis secun- 
dariis in dimidio quoque, in foliis vetustis siccis nervis 
tertiis transversis conspicuis et reticulis oblongis 
brevibus formantibus, lamina ligulata supra olivacea 
infra pallide viridibus et glaucis, basi amplexicauli 
inermi sed ex 10 cm marginibus cum aculeis 1—2.5 
mm longis 2-5 mm separatis subulatis rectis vel 
arcuatis albis, midnervo infra ex 19 cm cum aculeis 
1 mm longis 3-10 mm separatis subulatis adpresse 
reflexis fragilibus, in sectione mediali marginibus cum 
aculeis 1-2 mm longis 1-2.5 mm separatis crasse 
subulatis albis valde adscendentibus, midnervo infra 
cum aculeis 0.5-1 mm longis 2-4 mm separatis 
subulatis fragilibus albis adpresse adscendentibus, in 
