Page 425: Revision of Pandanus, 27 — St. John 
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subulatis compressis, in apice marginibus cum 
dentibus 0.7-2. 5 mm longis 1-4 mm separatis irregu- 
laribus biformatis majoribus latiter arcuato-subulatis 
adscendentibus albis cum apicibus minime brunneis 
olim bidentatis, midnervo infra cum subulato-serrulis 
0.5-1 mm longis, inflorescentia foeminea terminali 
erecta spicata cum 9 syncarpiis in 3 seriebus, pedun- 
culo 30 cm longo in apice 5 cm diametro clavato 
3-laterato cum 2 bracteis basilaribus foliaceis 80 cm 
longis 10 cm latis et cum bracteis binis caulinaribus 
caducis, rhachide 18 cm longo, syncarpiis congregatis 
12-15.5 cm longis 8.5-9 cm latis 6-7.5 cm crassis 
ovoideo-cylindricis sed latere proximo piano cum 
65-71 phalangibus in seriebus verticalibus, axili 
carnoso et fibroso, phalangibus 3-3.5 cm longis 2-3.1 
cm latis 1.7-2. 6 cm crassis aurantiaco-luteis obovoi- 
deis sed basi longiter cuneato-obpyramidali truncato 
parte *4 supera libera pyramidali-hemispherico (5)- 
6-anguloso apice truncato vel vadoso concavo 7-10 
mm lato orbiculari vel elliptico et cum ( 3-4) -5-7- 
(8) stigmatibus in margine circuli (rare cum stigmate 
unico centrali), stigmatibus 1.5-2 mm longis brun- 
neis papillosis horizontalibus superfkialibus cordatis 
vel reniformibus vel cordato-ovatis plerumque cen- 
trifugalibus sed paucis tangentialibus enim paucis 
centripetalibus, sinibus 1 mm longis profundis, endo- 
carpio mediali osseoso brunneo lateribus lateralibus 
3- 4 mm crassis intra laevibus et lucidis, seminibus 
6—8 mm longis late ellipsoideis, mesocarpio apicali 
medulloso et cum fibris longitudinalibus, mesocarpio 
basali fibroso et carnoso. 
diagnosis of holotype: Trees to 10 m 
tall, 20 cm in diameter; crown wide; bark 
brown, with numerous, sharp conic spines; 
branchlets 8.5 cm in diameter; leaf scars 6 mm 
high; prop roots none or less commonly present 
and up to 1 m long and 2.5 cm in diameter, 
light brown, with small, remote, short conic 
spines; leaves 1.58-1.65 m long, 9-11.6 cm 
wide, thick coriaceous, gently down-curved but 
not breaking across, dull, above olive-green, 
below green, narrowly furrowed above the mid- 
rib, in section V-shaped but somewhat curving 
towards the margin, not pleated, at midsection 
with 58 parallel secondary veins in each half, 
no visible tertiary cross veins except at the very 
base, blade ligulate-sword-shaped, narrowing to 
a stout, trigonous, subulate apex, this at 10 cm 
down 6-14 mm wide, the base amplexicaul, 
unarmed, white, and with membranous, entire, 
lateral flanges 1-1.5 cm wide and 16-20 cm 
long, at base beginning at 15—20 cm the spines 
4- 10 mm apart, ascending, deltoid with subu- 
late brown tips, otherwise opaque white, of two 
sizes, the major ones 6-8 mm long, the smaller 
ones 3-5 mm long; the midrib below beginning 
at 25 cm with spines 3-4 mm long, 3-12 mm 
apart, the first several reflexed subulate from 
broad conic bases, the later ones ascending; at 
midsection the margins with spines 3-5 mm 
long, 4-12 mm apart, irregular, large or small, 
broad subulate, ascending; the nearby midrib 
below with prickles 2.5-3 mm long, 6-15 mm 
apart, flat arcuate subulate, irregular; on the 
subulate apex the margins with teeth 0.7-2. 5 
mm long, 1-4 mm apart, irregular, the smaller 
straight subulate and salient, white except for 
the slightly brown tip, the larger ones broad 
arcuate subulate, ascending, occasionally biden- 
tate; the midrib below with prickles 0.5-1 mm 
long, 2-5 mm apart, subulate-serrations; pistil- 
late inflorescence terminal, erect, spicate, bear- 
ing 9 syncarps attached in three rows ; peduncle 
30 cm long, 5 cm in diameter, clavate, in section 
deltoid but the angles obtuse, subtended by two 
basal, persistent foliaceous bracts 80 cm long, 
10 cm wide, and the stalk bearing two caducous 
bracts; rhachis 18 cm long; syncarps crowded, 
12-15.5 cm long, 8.5-9 cm broad, 6-7.5 cm 
thick, in profile ovoid-cylindric but the proximal 
face flat or nearly so, bearing 65-71 phalanges 
in nearly vertical rows; core fleshy and fibrous; 
phalanges 3-3.5 cm long, 2-3-1 cm wide, 1.7- 
2.6 cm thick, orange-yellow, obovoid but the 
base long cuneate obpyramidal, truncate, the 
upper y 3 free, pyramidal-hemispheric, ( 5 ) — 6- 
angled, the apical area truncate or shallow con- 
cave, 7-10 mm wide, circular or elliptic, (3-4)- 
5-7-(8) -celled, very rarely with one central 
stigma, normally with all on the margins of the 
apical circle or polygon ; stigmas 1.5-2 mm long, 
brown, papillose, flush, horizontal, cordate to 
reniform to cordate-ovate, mostly centrifugal, 
but some tangential, and a few centripetal, pro- 
longed beyond the central crease is a deep sinus 
about 1 mm long; endocarp median, bony, 
brown, the lateral walls 3-4 mm thick, the inner 
surface smooth, shining; seeds 6-8 mm long, 
broad ellipsoid; apical mesocarp of solid pith, 
traversed by longitudinal fibers; basal mesocarp 
fibrous and fleshy. 
holotypus: Madagascar, lie Sainte Marie, 
Lambountsira, 4 km S of Ambodifototra, forest 
remnant in cultivated fields, 4 m alt, Aug. 26, 
1961, H. St. John 26,553 (bish). 
