126 
PACIFIC SCIENCE, Vol. XXII, January 1968 
section Dauphinensia, as is its closest relative, 
Pandanus centrzfugalzs St . John, from lie Ste. 
Marie, a species which has the phalanges 
( 3-4) -5-7-(8) -celled, 3-3.5 cm long, 
2-3.1 cm wide; stigma 1.5-2 mm long; endo- 
carp with lateral walls 3-4 mm thick ; tree with 
wide crown; leaves 1.58-1.65 m long, 9-11.6 cm 
wide, at midsection with 58 parallel secondary 
veins in each half, and the margin beginning 
15-20 cm from the base with spines 6-8 mm 
long. P. erectus has the phalanges (2) -3-4- 
celled, 2.6-3. 1 cm long, 1.5-2. 2 cm wide; 
stigmas 1-1.5 mm long; endocarp with lateral 
walls 1 mm thick; tree with the few branches 
strictly ascending; leaves 3.2-3.25 m long, 
15-16 cm wide, at midsection with 83 parallel 
secondary veins in each half, and the margins 
beginning 38 cm from the base with prickles 
2-2.5 mm long. 
In P. erectus but few of the phalanges fall to 
the ground, as most are caught and held in 
the leaf axils. Some germinate there, and may 
fall to the ground when the holding leaf falls. 
Several examples of this were observed on the 
collection St. John 26,566. 
The new epithet is the Latin adjective erectus, 
erect, and is given in reference to the strictly 
ascending branches. 
Pandanus spicatus sp. nov. (sect. Dauphinen- 
sia ) 
Figs. 260, 261 top right and bottom left 
nom. vern.: "vacoua madik” (= Pandanus 
small) . 
diagnosis holotypi: Arbor 8 m alta 15 cm di- 
ametro cum ramis pluribus suberectis eis in apice 
15 cm diametro et cum restis marcescentibus foliarum 
obtectis, cortice pallide brunneo cum spinis multis 
late conicis, radicibus fulturosis nullis, foliis 1.6-3. 2 
m longis 15-16.5 cm latis ligulatis crassiter coriaceis 
sublucidis supra obscure olivaceis infra olivaceis supra 
midnervum late U-sulcatis 2-plicatis in sectione me- 
diali cum 96-97 nervis parallelis secundariis in 
dimidio quoque nervis tertiis nullis lamina in apice 
valido trigono subulato subiter contracto eo 10 cm 
ex apice 18 mm lato basi amplexicauli inermi rosacea, 
marginibus ex 21-26 cm cum spinis 3-5 mm longis 
4-16 mm separatis valide subulatis ex basi conica 
lata adscendentibus infimis rosaceis alteris subviridibus, 
midnervo infra ad medium vel ultra inermi, in 
sectione mediali marginibus cum spinis 3-3.5 mm 
longis 8-26 mm separatis valide arcuato-subulatis 
compressis valde adscendentibus viridibus cum apice 
breve brunneo et olim margine uno cum spinis brevi- 
oribus vel enim in parte inermi, midnervo infra in 
sectione mediali vel distancia parva ultra cum aculeis 
1.5-2 mm longis 7-14 mm separatis subulatis sub- 
viridibus proxime adscendentibus, in apice marginibus 
et midnervo infra cum subulato-serris 0.7-1 mm longis 
2- 5 mm separatis, infructescentia terminali erecta cum 
circa 9 syncarpiis in fascicula foliarum clausis et paene 
drupis omnibus retentis, pedunculo 20 cm longo 2 cm 
diametro carnoso, syncarpiis circa 9 cm longis et 7 cm 
diametro ovoideis compressis cum drupis multis, eis 
20-26 mm longis 10-20 mm latis 10-17 mm crassis 
(sed in geminis 27 mm latis et 6-7-loculatis) licet 
luteis cuneatis vel obovoideis 3-5-loculatis parte I 
supera libera laevibus lateribus contiguis subplanis, 
stigmatibus 2-2.5 mm latis cordatis horizontalibus 
exprojectentibus brunneis papillosis circa apicem con- 
gregate, inter stigmates licet cum valleculis vadosis, 
endocarpio centrali 10 mm longo pallide brunneo 
osseoso lateribus lateralibus 1.5-2 mm crassis intra 
transverse rugosis et cum projectione apicali in locula 
omnina, mesocarpio apical i cum fibris longitudinalibus 
fortibus, mesocarpio basali fibroso et carnoso. 
DIAGNOSIS OF holotype : Tree 8 m tall, 15 
cm in diameter, with several suberect branches, 
these at apex 15 cm in diameter and long 
clothed with persistent leaf bases; bark light 
brown, with numerous broad conic spines ; prop 
roots none; leaves 1.6-3. 2 m long, 15-16.5 cm 
wide, ligulate, thick coriaceous, somewhat shiny, 
above dark olive-green, below olive-green, 
with a broad U-shaped furrow above the midrib 
and two lateral pleats, at midsection with 96-97 
parallel secondary veins in each half, no visible 
tertiary cross veins, the blade abruptly narrowed 
to a stout trigonous subulate apex, this 10 cm 
down 18 mm wide, the base amplexicaul, un- 
armed, pink, but beginning 21-26 cm up the 
margins with spines 3-5 mm long, 4-16 mm 
apart, stout subulate, ascending from a broad 
conic base, the lowest pink, the next greenish, 
the midrib below unarmed to or beyond the 
middle; at midsection the margins with spines 
3- 3.5 mm long, 8-26 mm apart, heavy arcuate 
subulate, flattened, strongly ascending, green, 
with short brown tips, and not infrequently 
one of the margins has shorter spines or even in 
places is unarmed for a short span; the midrib 
below, beginning at or shortly above the middle, 
with prickles 1.5-2 mm long, 7-14 mm apart, 
subulate, closely ascending, greenish; near the 
apex the margins and midrib below with 
subulate-serrations 0.7-1 mm long, 2-5 mm 
apart; infructescence terminal, erect, bearing 
about 9 syncarps, concealed in the large plume 
of erect or ascending leaves which trap the 
! 
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