Page 439: Revision of Pandanus, 27 — St. John 
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drupes but few of which fall to the ground 
when shed; peduncle 20 cm long, 2 cm in 
diameter, fleshy; syncarps about 9 cm long and 
7 cm in diameter, ovoid, compressed, bearing 
numerous drupes, these 20-26 mm long, 
10-20 mm wide, 10-17 mm thick (but in 
doubles 27 mm wide and with 6-7 cells), ap- 
parently yellow, cuneate to obovoid, 3-5 -celled, 
upper y 4 free, the surface smooth, the con- 
tiguous sides nearly flat; stigmas 2—2.5 mm 
wide, cordate, horizontal, flush, brown, papil- 
lose, approximate near the apex ; endocarp 
central, 10 mm long, pale brownish, bony, the 
lateral walls 1.5-2 mm thick, the inner surfaces 
transversely rugose, each cell bearing an 
acuminate projection toward the stigma; apical 
mesocarp with strong longitudinal fibers ; basal 
mesocarp fibrous and fleshy. 
holotypus: Malagasy Republika, Gravirage, 
7 km SW of Joffreville, Montagne d’Ambre, 
dense moist forest with "'rami” ( Canarium 
Boivini ), and "ambora” ( Tambourissa sp.), 
1,000 m alt, 14 Sept. 1961, H. St. John 26,577 
(bish). 
discussion: P. spicatus is a member of the 
section Dauphinensia, as is its closest relative, 
P. dauphinensis Martelli in Martelli & Pic. Ser., 
of coastal southeastern Madagascar, a species 
with the peduncle 46 cm long, 5.5 cm in diam- 
eter; syncarps 17.5-21 cm long, cylindric; 
phalanges 16-23 mm wide, the upper y s free; 
endocarp smooth within; leaves at midsection 
with 70 parallel secondary veins in each half, 
the midrib below beginning at 40 cm up with 
spines 3-4 mm long, 7-12 mm apart, conic, 
acute, reflexed; at midsection the margins with 
spines of two kinds, the larger ones 3-5 mm 
long, and alternating with smaller ones 
2-2.8 mm long. P. spicatus has the peduncle 
20 cm long, 2 cm in diameter; syncarps about 
9 cm long, ovoid; phalanges 10-20 mm wide, 
the upper y 4 free ; endocarp rugose within ; 
leaves at midsection with 96-97 parallel secon- 
dary veins in each half, the midrib below un- 
armed to or beyond the middle; at midsection 
the margins with uniform spines 3-3.5 mm 
long. 
The new epithet is the Latin adjective, spica- 
tus, spike-bearing, and is given with reference 
to the spicate inflorescence. 
Section Mammillarisia 
Pandanus aridus sp. nov. (sect. Mammillarisia ) 
Figs. 262, 261 bottom right, 264 top left 
and right 
nom. vern.: "fandrana.” 
diagnosis holotypi: Frutex multiramosa, caulibus 
7 m altis 4 cm diametro, cortice griseo acute spinoso, 
radicibus fulturosis paucis 20-30 cm longis 2 cm 
diametro, foliis 35—45 cm longis 16—20 mm latis sub- 
coriaceis gradatim arcuatis supra obscure olivaceis 
infra pallide viridibus et glaucis supra midnervum 
anguste V-sulcatis deinde planis in sectione mediali 
cum 17 nervis parallelis secundariis nervis tertiis 
transversis nullis lamina ligulata sed dimidia ultima 
diminuenti in apice trigono subulato eo 10 cm ex 
apice 6.5 mm lato basi amplexicauli et inermi alba sed 
ex 1.3-2 cm marginibus cum aculeis 2-2.3 mm longis 
2-12 mm separatis deltoideo-subulatis adscendentibus 
albis sed apicibus brunneis, midnervo infra ex 3.5-4 
cm cum aculeis 1.3-2 mm longis 4-11 mm separatis 
albis subulatis valde reflexis, in sectione mediali 
marginibus cum aculeis 0. 5-0.9 mm longis 7-13 mm 
separatis gracilibus subulatis divergentibus albis, in 
apice subulato marginibus et midnervo infra cum 
subulato-serrulis 0. 3-0.7 mm longis 1.5-4 mm sepa- 
ratis, inflorescentia foeminea terminali erecta cum 
syncarpio unico, pedunculo 8 cm longo 8 mm diam- 
etro 3-laterato folioso-bracteato et in apice cum 6 
bracteis marcescentibus exteriori 2.5 cm longa 1 cm 
lata elliptica naviculari marginibus spinoso-fimbriatis, 
syncarpio 5.5-6 cm diametro globoso viridi cum 21 
phalangibus eis 22-25 mm longis 12-23 mm latis 
10—22 mm crassis obovoideis firme connatis parte i 
supera libera subpyramidali-conica 5-6-angulosis 2-4- 
loculatis apice lobato sinibus 0.5 mm profundis, stig- 
matibus 1-1.5 mm latis cordatis obscure brunneis 
papillosis centripetalibus horizontalibus minime ele- 
vatis congregatis, endocarpio 13-14 mm longo supra- 
mediali brunneo osseoso solido obovoideo lateribus 
lateralibus 2 mm crassis intra pallidis laevibus lucidis, 
seminibus 6—8 mm longis ellipsoideis, mesocarpio 
apicali parvo medulloso et cum fibris paucis, meso- 
carpio basali carnoso et fibroso. 
DIAGNOSIS of holotype: Shrub with many 
stems, 7 m tall, 4 cm in diameter; bark gray, 
sharp spiny; prop roots few, 20-30 cm long, 
2 cm in diameter; leaves 35-45 cm long, 
16-20 mm wide, subcoriaceous, gently curving, 
above dark olive-green, below pale green and 
glaucous, above the midrib with a narrow 
V-shaped furrow, otherwise nearly plane, at 
midsection with 17 parallel secondary veins in 
each half, no tertiary cross veins, blade ligulate 
but the outer half tapering to a trigonous, 
subulate apex, this 10 cm down 6.5 mm wide, 
the base amplexicaul, unarmed, white, but begin- 
