Page 441 : Revision of Pandanus, 27 — St. John 
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ning at 1.3-2 cm the margins with prickles 
2-2.3 mm long, 2-12 mm apart, deltoid subu- 
late, ascending, white with brownish tips ; the 
midrib below beginning at 3.5-4 cm with 
prickles 1.3-2 mm long, 4-11 mm apart, subu- 
late, reflexed, close, white; at midsection the 
margins with prickles 1—1.8 mm long, 6-15 mm 
apart, subulate, arcuate ascending, white; the 
midrib below with prickles 0. 5-0.9 mm long, 
7-13 mm apart, slender subulate, divergent, 
white; on the subulate tip the margins and 
midrib below with subulate serrulations 0.3- 
O. 7 mm long, 1.5-4 mm apart; pistillate in- 
florescence terminal, erect, bearing a solitary 
syncarp; peduncle 8 cm long, 8 mm in diam- 
eter, 3 -sided, leafy bracted, and at the apex 
with 6 marcescent bracts, the outer one 2.5 cm 
long, 1 cm wide, elliptic, navicular, the margins 
spiny fimbriate; syncarp 5.5-6 cm in diameter, 
globose, green, bearing 21 phalanges, these 
22-25 mm long, 12-23 mm wide, 10-22 mm 
thick, obovoid, firmly connate, upper y 3 free, 
slightly pyramidal conic, 5-6-angled, 2-4-celled, 
lobed about 0.5 mm deep between the apices 
of the carpels; stigmas 1-1.5 mm wide, cordate, 
dark brown, papillose, centripetal, horizontal, 
slightly raised, congregate; endocarp 13-14 mm 
long, supramedian, brownish, bony, massive, 
obovoid, the lateral walls 2 mm thick, the inner 
surfaces pale, smooth, shining; seeds 6-8 mm 
long, ellipsoid; apical mesocarp scant, of pith 
and a few fibers; basal mesocarp fleshy and 
fibrous. 
holotypus: Malagasy Republika, Rano- 
mainty, 18 km E of Amboasary, W of Fort- 
Dauphin, dry arid scrub with Euphorbia sp., 
Croton sp., 175 m alt, Sept. 26, 1961, H. St. 
John 26,382 (bish). 
discussion: P. aridus is a member of the 
section Mammillarisia, as is its closest relative, 
P. saxatilis Martelli, of northern Madagascar, 
a species with the phalanges 23-27 mm long, 
the upper y 2 free; endocarp occupying almost 
the whole interior of the phalange; lower bract 
of the inflorescence 2 cm wide ; prop roots none. 
P. aridus has the phalanges 22-25 mm long, 
the upper y 3 free; endocarp 13-14 mm long, 
slightly supramedian, occupying about % of the 
interior of the phalange; basal mesocarp 7 mm 
long; lower bract of the inflorescence 1 cm 
wide; prop roots few, 20-30 cm long, 2 cm in 
diameter. 
The new epithet is the Latin adjective, aridus, 
dry, and is given with reference to the dry forest 
habitat. 
Pandanus connatus sp. nov. (sect. Mammillari- 
sia) 
P. acuminatus Hort. ex Wendl., Ind. Palm. 
45, 1854, nomen nudum. 
Vinsonia acuminata Gaud, ex Warb., Engler’s 
Pflanzenreich IV,9:89, 1900, nomen 
nudum, published in synonymy. 
Figs. 263, 264 bottom left and right 
nom. vern. : "vacoua madik” (= Pandanus 
small) . 
diagnosis holotypi: Arboriformata 10 m alta 9 
cm diametro, cortice brunneo cum spinis conicis brevi- 
bus, ramulis 2.5 cm diametro, radicibus fulturosis 
nullis, foliis 90-109 cm longis 2-2.3 cm latis curvatis 
sed non fractis ligulatis subcoriaceis supra obscure 
olivaceis infra intense viridibus supra midnervum sil- 
lonatis V-formatis et in media ultima 2-plicatis in sec- 
tione mediali cum 23-27 nervis parallelis secundariis 
in dimidio quoque nervis tertiis solum in basi in 
tertia ultima sensim in apice trigono subulato diminu- 
entibus eo in puncto 10 cm ex apice 6 mm lato basi 
amplexicauli inermi sed ex 6-7.5 cm marginibus cum 
aculeis 1.5-2. 5 mm longis 3-8 mm separatis crassiter 
arcuato-subulatis adscendentibus albis cum apicibus 
brunneis, midnervo infra per I partem inermi, in sec- 
tione mediali marginibus cum serrulis 0. 3-0.9 mm 
longis 3-7 mm separatis albis cum apicibus brunneis, 
in apice marginibus et midnervo infra cum serrulis 
0.2-0. 5 mm longis 1-3.5 mm separatis albis cum 
apicibus brunneis, inflorescentia foeminea erecta 
terminali, pedunculo 8 cm longo in apice 2 cm diam- 
etro clavato trigono folioso-bracteato, bracteis superis 
4-5 cm longis 2.5-3 cm latis deltoideis subulatis 
ciliatis papyraceis quando siccis brunneis, syncarpio 
solitario 9.5 cm diametro globoso cum 79 phalangibus 
eis 3. 3-3. 5 cm longis 1. 6-2.1 cm latis 1.3-1. 7 cm 
crassis connatis oblanceoloideis compressis 1-3-locu- 
latis 5-6-angulosis laevibus lucidis parte 3 supera 
libera lateribus liberis subcurvatis illis contiguis 
planis, apicibus cum 2-3 stigmatibus cum vallecula 
0.5 mm profunda subclausa, stigmatibus 1.5 mm 
diametro asymmetricalibus cordatis horizontalibus 
superficialibus pallide brunneis papillosis, endocarpio 
20-23 mm longo supramediali osseoso brunneo cum 
humeris medialibus latis lateribus 5-6 mm crassis 
intra laevibus et lucidis, seminibus 10-12 mm longis 
lineari-ellipsoideis, mesocarpio apicali medulloso et 
cum fibris paucis, mesocarpio basali fibroso et carnoso. 
diagnosis of holotype : Tree-like, 10 m 
tall, 9 cm in diameter; bark brown, with short 
conic spines; branchlets 2.5 cm in diameter; 
prop roots none; leaves arching and recurving, 
