416 
PACIFIC SCIENCE, Vol. XXII, July 1968 
anthers 4-5 mm long, 0.5 mm wide, almost 
linear, at apex bearing a subulate prolongation 
of the connective 0. 3-0.4 mm long. 
lectotype: Iter Australiense, 1802-5, R- 
Brown 5,199, the pistillate tessellate part (bm), 
but, as his manuscript reveals, this is more pre- 
cisely: Gulf of Carpentaria, [Queensland, Wel- 
lesley Group of Islands], Island C [Allen L, 
Island Bay], Nov. 22, 1802, R. Brown 5,799, 
but only the two phalanges with broad tessellate 
apices (bm). 
specimens examined: Australia, same data, 
the staminate inflorescence, R. Brown 5,799 
(bm, k). 
discussion: P. spiralis is a member of the 
section Eydouxia. It was one of the first two 
species discovered in Australia. Though remark- 
ably distinct, it has had varied treatment, per- 
haps due to the brevity of the original descrip- 
tion. Solms-Laubach (Linnaea 42:79, 1878) 
listed it with the Species Dubiae. Balfour (Linn. 
Soc, J. Bot. 17:56, 1880=1878) made it a 
synonym of P. odoratissimus L. f., as did War- 
burg (Pflanzenreich IV, 9:46, 1900, and Ben- 
tham & von Mueller (FI. Austral. 7:149, 1878). 
It was reestablished by Martelli (Webbia 
4(1) :31, 90, 94, 1913; 4(2) :t. 13, fig. 1-2, 
1914). 
Various more recent collections from more 
or less distant localities have been identified as 
P. spiralis, but all, it seems, incorrectly so. S. T. 
Blake (Austral. J. Bot. 2:130, 1954) stated 
that Martelli’s published illustration was drawn 
from Hulse’s collection from Escape Cliffs. That 
was implied by the wording of Martelli’s 
legend: "P. spiralis: Australia, Cliff. (H. M. 
ex H. Brit. Mus.).” However, the specimen 
referred to: Northern Territory, Escape Cliffs, 
Hulls [an error for W . Hulse], (mel), has the 
phalanges 6. 6-7. 9 cm wide, has only 8-9 
carpels, the central apical sinuses 1.5-3 mm 
deep, the apical mesocarp cavities 8-11 mm 
long, suborbicular; and the stigmas 4-5 mm 
long or wide. This specimen is now determined 
as P. convexus St. John (Pacif. Sci. 15:182-184, 
fig. 10, 1961) and is easily distinguishable from 
P. spiralis. Martelli’s illustration of P. spiralis 
does not agree with the Hulse collection of P . 
convexus, but on the other hand it does exactly 
agree with the broad, many carpeled, unlobed 
phalanges that are the lectotype of P. spiralis 
R. Br., and it is evident that Martelli really 
made the drawing from Brown’s type. Appar- 
ently there are no subsequent collections of 
Pandanus from the Wellesley Islands in the 
Gulf of Carpentaria. 
Martelli’s illustration of P. spiralis (Webbia 
4(1) : t. 13, fig. 1-2, 1913) showed as Figure 
1 an oblique lateral view of a phalange exactly 
like the larger of the two still in the British 
Museum of Natural History. His Figure 2 
shows a median longitudinal section of a similar 
phalange. This halved phalange was not found 
in 1963 in the London herbarium material. 
Brown’s collection of fruiting P. spiralis, still 
in the British Museum, is a mixture. Besides the 
lectotype, discussed above, it also contained 
fruits of a very different species which will be 
described as P. Brownii St. John. 
Section Pandanus 
Pandanus Brownii sp. nov. (sect. Pandanus ) 
Fig. 270 
diagnosis holotypi: Solum 2 phalanges, eis 
5.9-6. 2 cm longis 7. 2-8.8 cm latis 4. 5-4.7 cm 
crassis anguste vel late flabellatis compressis cum 6 
angulis majoribus et carpelis omnibus in fuga salienti- 
bus et aliquis cum 2-3 fugis longitudinalibus brevibus 
et valleculis solum in medio lateribus laevibus lucidis 
curvatis parte l / 2 supera libera apice rotundato suturis 
lateralibus omnibus profundis et prominentibus et 
aliquis in basi extends, sinibus apicalibus centralibus 
3-4 mm profundis in centra V-formatis subrectis 
deinde recurvatis, carpelis 19-29 plerumque in 3 
seriebus eis centralibus oblate pyramidali-semiorbicu- 
laribus illis marginalibus cum stigmatibus in margine 
apparentibus et cum regione lato concavo subverticali 
distali, sinibus proximalibus V>-% a d fondam ex- 
tends, stigmatibus 2-3 mm latis reniformibus plerum- 
que prostratis eis centralibus horizontalibus illis 
marginalibus extraobtutis, endocarpio solido 4 cm 
longo paene submediali osseoso obscure mahogani- 
colorato lateribus lateralibus 10-11 mm crassis intra 
circum-striatis lucidis, seminibus 15-18 mm longis 
ellipsoideis, mesocarpio apicali medulloso denso cum 
fibris longitudinalibus, sed non in sectionibus partitis, 
mesocarpio basali parvo fibroso et carnoso. 
DIAGNOSIS OF HOLOTYPE: It consists of 2 
phalanges only; phalanges 5. 9-6.2 cm long, 
7. 2-8. 8 cm wide, 4. 5-4. 7 cm thick, narrowly 
to broadly fan-shaped in outline, compressed, 
with 6 major angles and each carpel salient and 
