418 
forming another ridge and some of the carpels 
with 2-3 short longitudinal ridges and valleys 
only at midsection, the sides smooth, shining, 
curving, upper y 2 free, apex rounded, all lateral 
sutures deep and prominent and several of them 
extending to the very base; central apical sinuses 
3-4 mm deep, V-shaped at center, then re- 
curving, nearly straight; carpels 19-29, mostly 
in 3 rows, the central ones oblate pyramidal- 
semiorbicular, the marginal ones with the stig- 
mas appearing marginal and with a broad con- 
cave, nearly vertical surface where the outer side 
reaches the stigma, and a few of the carpels 
with the stigma slightly remote from the margin 
which bears a small distal concavity; proximal 
sinus running y 2 — % way to valley bottom; 
stigmas 2-3 mm wide, reniform, mostly flush, 
the central ones horizontal, the marginal ones 
flush but facing outward; endocarp massive, 4 
cm long, slightly submedian, bony, dark ma- 
hogany-colored, the lateral walls 10-11 mm 
thick, the inner surfaces striate ringed, shining; 
seeds 15—18 mm long, ellipsoid; apical meso- 
carp of dense pith, traversed by longitudinal 
fibers, but not forming perceptible carpelary 
cavities or structures; basal mesocarp scant, 
fibrous and fleshy. 
holotypus: Australia, Iter Australiense, 
1802-5, R. Brown (bm). 
discussion: P. Brownii is a member of the 
section Pandanus, as is the most similar species, 
P. Delessertii Warb., a species known only from 
the type collection, published by Gaudichaud 
without description as Eydouxia Delessertii and 
said to come from Bourbon Island. No one 
has found it there since, and the detailed and 
careful monograph of the Mascarene species by 
Vaughan and Wiehe discredits it, and leads us 
to believe that the species is native elsewhere. 
It has 19 carpels in 3 rows in the fan-shaped 
phalange, 4.2 cm long, 6.5 cm wide, 4 cm 
thick, compressed, flabellate in outline; central 
apical sinuses little more than creases 0-1 mm 
deep; stigmas 4-6 mm long, cordate shield- 
shaped, and all that still remain are obtuse 
(none acute as shown in Gaudichaud’s pi. 18, 
fig. 7-8). The holotype (p) consists of a single 
phalange. P. Brownii differs in having the 
phalanges of 19-29 carpels, 5. 9-6. 2 cm long, 
7. 2-8. 8 cm wide, 4. 5-4. 7 cm thick; central 
PACIFIC SCIENCE, Vol. XXII, July 1968 
apical sinuses 3-4 mm deep, V-shaped; stigmas 
2-3 mm wide, reniform. Species of this general 
type are known only from Australia; so it is 
concluded that P. Delessertii Warb. is probably 
an Australian species of which Gaudichaud 
obtained the specimen from some other botanist. 
The holotype of P. Brownii was mixed with 
the lectotype of P. spiralis R. Br., from Allen 
Island, Wellesley Group, Gulf of Carpentaria, 
Queensland, Australia. The type collections of 
both of Brown’s two new species, P. spiralis 
and P. pedunculatus, contain mixtures of other 
species. It appears that either he or his suc- 
cessors who accessioned his herbarium con- 
sidered each a variable species and added to the 
boxes other loose fruits which they considered 
similar. There is no implication that this new 
species and P. spiralis were collected at the same 
locality. In his manuscript (British Museum) 
he also has a description of a species from 
Endeavour Straits, which he called P. odoratis- 
simus L. f., but no separate specimen of this is 
now found in his herbarium in London. There 
is such a specimen of Brown’s collecting now 
in Edinburgh. It is labeled " Pandanus odoratis- 
simus ? North Coast.” It is a good staminate 
inflorescence of a species in the section Pan- 
danus, but it cannot be more precisely identified. 
Pandanus pedunculatus R. Br., Prodr. FI. Nov. 
Holl. 341, 1810 (sect. Pandanus ). 
Fig. 271 
EXPANDED DIAGNOSIS OF LECTOTYPE: "Tree 
4-5.3 m tall; trunk erect, 1.3-2. 6 m tall, below 
the middle bearing at an acute angle, prop roots 
that are terete, roughened by remote, small 
tubercles; branches divaricate, sometimes re- 
clining; leaves in terminal, loose fascicles, broad 
sword-shaped, above green, below glaucous, 
striate with fine but prominent veins, the mar- 
gins ascending denticulate almost to the base, 
the keel retrorsely roughened scarcely up to the 
point 1/3 from the apex; syncarp solitary, termi- 
nal, pedunculate, nodding, ovate-globose, of the 
size of a human head or somewhat more elon- 
gate, bearing numerous compact phalanges,” 
(this much extracted from Brown’s manu- 
script) ; phalange 7.6 cm long, 4.7 cm wide, 
3.9 cm thick, pyriform, 4-angled, the sides 
gently convex, smooth, shining, of the 7 lateral 
sutures 5 are sealed but 2 are narrow and 
